Charles Griswold

  • Curator and Schlinger Chair of Arachnology
  • Entomology
  • PhD

World taxonomic revision and phylogeny of Goblin spiders.


 

 

In the past 150 years 500 species of goblin spiders have been described: we estimate that the true number is greater than 2000.  We are attempting to compress several lifetimes worth of work into 7 years by combining and coordinating the efforts of 40 researchers from around the world, utilizing on-line descriptive and phylogenetic databases, and making extensive use of digital imaging technology.  In essence we are attempting to reinvent taxonomy so that this may be done rapidly, consistently, comprehensively, and producing user-friendly products that may be accessed by anyone with access to the internet.  CAS participants include Darrell Ubick and Anthea Carmichael (technicians), graduate students Alma Saucedo (San Francisco State) and Natalia Chousou Polydouri (UC Berkeley), former postdoctoral researcher Fernando Álvarez Padilla and myself.  This research is supported in large part by the Goblin Spider PBI NSF grant.

Goblin Spider Website

 


 

Evolution of spiders


 

 

 

 

 

 

 

 

 

 

 

Spider phylogeny has been revolutionized in the last 40 years.  Nevertheless, the problem of estimating the phylogeny of all spiders is beyond the grasp of any one person.  The Assembling the Tree of Life (AToL) Spiders project brings together the talents of more then 20 researchers from 12 countries to organize existing and develop new data from morphology, behavior and molecules to produce a robust phylogeny for all spider taxa of the tribal level and above.  We have tissues for more than 1200 species, have a morphological matrix of more than 900 characters, and have developed an ontology of spider morphology.  My role has been particularly as a mentor (at CAS three SSI undergraduates, five graduate students, and three postdoctoral researchers have learned to contribute data for this project), as a collector of specimens and tissues (grant money helped support expeditions to Australia, Myanmar, Madagascar, New Zealand and South Africa), and as an expert on spider morphology.

ATOL: Phylogeny of Spiders Website

 


 

Survey of biotic hotspots in Asia and Africa



 

 

 

 

 

 

 

 

 

 

 

I am involved in two surveys of biotic hotspots (Madagascar and Gaoligongshan, China).  Both have been supported by grants from the NSF BSI program.  Several expeditions by myself and students have produced more than a million specimens, and more than 750,000 specimens are on loan from the Madagascar project alone.  For the Madagascar project I mentored a postdoctoral researcher (Silva), two SFSU graduate students (Andriamalala and Wood) and five Lakeside interns and have so far contributed to nine publications.  For the Gaoligongshan project I have mentored two postdoctoral fellows (Wang and Miller) and two Lakeside fellows, and have so far contributed to 15 publications.  Graduate student Wood, curatorial assistant Carmichael and I are completing a revision and molecular phylogeny of the spider family Phyxelididae in Madagascar, and graduate student Joel Ledford and I am working on a revision of the spider family Zoropsidae in Madagascar (about 70 new species).  With current and former postdoctoral fellows Miller and Wang, I am preparing major revisions of Gaoligongshan symphytognathoid (micro-orb weavers) and coelotine (dark forest runner) spiders.

Gaoligongshan Website

Madagascar Website

 


 

Comparative morphology of spiders


 

 

 

 

 

 

 

 

 

 

New (e.g., the Scanning Electron Microscope, SEM; digital imaging) and old (e.g., microtome) tools have enabled us to study the morphology of spiders in greater detail than ever before, worldwide collections such as at CAS allow us to compare features across all spiders, and the internet enables us to share and compare these data globally.  Several research projects ongoing in the CAS Arachnology lab, including student projects on assassin spiders and cave spiders, the Goblin spider PBI, and the Spiders AToL project contribute to a broad documentation of spider morphology, which in turn poses and informs questions on the evolution and function of these morphologies.

Lina Almeida

I am a PhD student in Brazil at the Laboratório de Artropodes of Instituto Butantan and at Universidade de São Paulo.  I was a Lakeside Scholar at CAS during April—June 2009.  I am interested in the systematics and evolution of spiders, especially the families Titanoecidae and Amaurobiidae, and for my PhD I am studying the amaurobiid subfamily Macrobuninae.

linamas@gmail.com


Fernando Alvarez-Padilla

I am a Professor at Universidad Nacional Autónoma de México.  I got my PhD degree at George Washington University and was a postdoctoral researcher at CAS from 2007-2009.  I am interested in the systematics, evolution and comparative morphology of spiders, especially the Tetragnathidae and Oonopidae.  I am a participant in the Goblin spiders Planetary Biodiversity Inventory (PBI).  My research at CAS took me on expeditions to Madagascar and Mexico.

fap@ciencias.unam.mx


Daniela Andriamalala

I am a PhD student at George Washington University doing research for the Goblin spiders Planetary Biodiversity Inventory (PBI).  My main research interests are in phylogeny, evolution and biogeography of spiders: relationships between ecologically linked taxa, divergence time estimation using molecular sequence data and phenomenona related to changes in rates of evolution in different spiders groups.  For my PhD thesis research, I am investigating the systematic, evolution and biogeography of the goblin spider genus Lionneta (Oonopidae), endemic to Madagascar and the Seychelles Islands.  I am also interested in the relationships between the genera within the Oonopid group to which they belong.  At CAS (2005—2007) I did a MSc thesis at San Francisco State University on the systematics and evolution of the Madagascar jumping spider genus Padilla.

dand@gwmail.gwu.edu


Anthea Carmichael

I am a Curatorial Assistant in the Arachnology lab and a Molecular Lab Technician in the Center for Comparative Genomics.  I am in charge of preparing molecular data for spiders; assisting with the Goblin Spider PBI, and managing the arachnid and myriapods collections.  I completed a MSc at san Francisco State University studying the co-evolution of birds and their parasitic mites.

acarmichael@calacademy.org


Robin Carlson

I am a volunteer in the Arachnology lab.  I first became involved as an undergraduate intern in the CAS Summer Systematics Institute (SSI – 1996), studying the fine structure and function of spider silk and the ontogeny of spider silk spinning organs.  I am currently a manager with the Pacific States Marine Fisheries Commission and also a dancer and choreographer with the Blueprint Dance Company.


Natalia Chousou Polydouri

I am a PhD student at University of California, Berkeley. My current research is focused on evolution and biogeography of spiders of the IndoPacific region, especially the goblin spiders (Oonopidae) and Tetragnathidae. My research has taken me to Micronesia and to Brunei.

nataliacp@berkeley.edu


Joel Ledford

I am a PhD student at University of California, Berkeley.  I am broadly interested in the study of biodiversity, with a particular interest in the regional history and biogeography of California. I am a specialist on small, cryptic groups of spiders in Western North America.  As a participant in the CAS Summer Systematics Institute (SSI - 1997) I studied trap- door spiders (Migidae) in expeditions to Madagascar and Gulf of Guinea. My current research is focused on cave- adapted spiders in the families Leptonetidae and Telemidae. The goals of these projects include revisionary taxonomy and phylogenetics with the aim of developing an understanding of speciation in cave ecosystems.   Aside from spiders, I am a student of Aikido at River City Aikido in Sacramento, CA and an active member of Yolo Bowmen in Davis, CA where I practice field target archery.

Website

jledford@berkeley.edu


Giovannia Maki

I am an artist and scientific illustrator and have done many renderings for the Entomology Department, especially of spiders.  I have also created artwork for exhibits at California Academy of Sciences.  You can visit my portfolio online.

Website

gmaki@calacademy.org


Jeremy Miller

I am curator of Arachnida at the Naturalis Museum in Leiden, the Netherlands.  I was a postdoctoral researcher at CAS from 2005-2008, where I studied the evolution of black widow spiders (Latrodectus) and participated in the biodiversity inventory of the Gaoligongshan in Yunnan, China.  My research is focused on spider systematics, quantitative biodiversity, and bioinformatics. Observations from morphology, behavior, and molecular sequences all inform my work on spider systematics. I have described species belonging to a variety spider lineages and explored the evolution of key morphological and behavioral traits. I use specialized statistical techniques to estimate the number of species in a place. All but the most exhaustive inventories overlook species. Estimators driven by the number of rare species can provide correction factors. Emerging online resources have great potential for broadening dissemination of taxonomic information. Interactive tools offer new ways to simultaneously assist the research community and engage the public.

Website


Alma Saucedo

I am a masters student in the Arachnology Lab and at San Francisco State University and a trainee in the Goblin Spider PBI Project.  I have also collected specimens and data for the Spiders AToL Project.  I am studying the morphology, phylogeny and biogeography of goblin spiders of Madagascar (Oonopidae) and am revising some new goblin spider genera.  I participated in the 2009 CAS/Goblin spider PBI expedition to Madagascar.

asaucedo@calacademy.org


Diana Silva

I am curator of Arachnida at the Universidad de San Marcos in Lima, Peru.  I got my PhD degree from Cornell University and was a postdoctoral researcher at CAS from 2001 through 2005.  I contributed to the Madagascar BSI and to the Spiders AToL projects.  I am interested in the systematics and evolution of spiders, especially the tropical wolf spiders (Ctenidae) and giant crab spiders (Sparassidae).  My research at CAS took me on expeditions to Australia, French Guiana, Madagascar and New Zealand.

dianasil@gmail.com


Vic Smith

I am a Curatorial Assistant in the Arachnology lab and am the digital imaging specialist for the lab.  I also manage loans of Diptera (flies) and assist with management of the arachnid and myriapod collections.  I am currently imaging all the Diptera type specimens in the CAS collection.  I did my MSc at San Francisco State University studying sea slugs.

vsmith@calacademy.org


Warren Savary

I am a compliance officer at the U.S. Food and Drug Administration and Field Associate of the California Academy of Sciences.  My research interests include the systematics and phylogeny of the arachnid orders Solifugae and Scorpiones, and the biogeography of western North America. Since 2004, I have served as research collaborator and webmaster for the NSF-funded REVSYS Vaejovidae grant and, since 2007, as research collaborator and webmaster for the BS&I Solifugae grant.  (NSF-REVSYS Vaejovidae Grant, 2004–; NSF-BS&I Solifugae Grant, 2007–).

wsavary@yahoo.com


Tamas Szuts

I am the Schlinger Chair of Arachnology Postdoctoral Fellow and am studying the Salticidae (jumping spiders) of Madagascar, the phylogeny of hisponine jumping spiders, and am collaborating on studies of Malagasy zorocratid spiders.  After completing my Ph.D. on Salticidae at the University of Szeged in Hungary I served as a Carlsberg Postdoc and Marie Curie Postdoc at the Zoological Museum of the University of Copenhagen, where I studied the evolution of orb-weaving spiders.

tszuts@calacademy.org


Darrell Ubick

I am a curatorial assistant in the Arachnology Lab and am a collaborator with the Goblin Spider PBI Project, where I am revising several new genera of Oonopidae from Madagascar.  I also have long standing research interests in taxonomy and phylogeny of harvestmen (Opiliones) and in cave biology.  I am an editor and author for the Spider Guide for North America and have contributed to the Spiders AToL Project and biotic surveys of Madagascar and the Gaoligongshan, and which research has taken me on expeditions to China, Madagascar, Myanmar and South Africa.

dubick@calacademy.org


Hannah Wood

I am a doctoral student in the Arachnology Lab and at the University of California, Berkeley studying assassin spiders (Archaeidae and Mecysmaucheniidae) for my dissertation research. Using phylogenetics, my research seeks to understand biogeographical patterns on different land masses and how macro-speciation occurs. I am is also studying predatory behaviors and their associated biomechanical properties in order to understand the evolutionary trade-offs between different predatory strategies and am particularly interested in the interplay between behavior and morphology.  My research has taken me on expeditions to Australia, Chile, Madagascar, New Zealand and South Africa, where I have also collected specimens and data for the Spiders AToL Project and the Goblin Spider PBI Project.

hwood@berkeley.edu


XinPing Wang

I am a data manager and biostatistician at the University of Florida, and an arachnologist specializing in the spider subfamily Coelotinae.  I manage two websites, online Coelotinae (http://www.Amaurobiidae.com) and Endemic Spiders in China (http://www.ChineseSpecies.com).  I am interested in discovery, morphology, phylogeny, biogeography of coelotine and related spiders.  I was a postdoctoral researcher at the CAS Arachnology lab in 2000-2001, where my research focused on Coelotinae of the Gaoligong mountains in western China.

Education

University of California, Berkeley, Conservation of Natural Resources, B.Sc., 1972
University of California, Berkeley, Entomology, Ph.D., 1983
American Museum of Natural History, Entomology, Postdoctoral Fellow, 1987-1988
Smithsonian Institution, Entomology, Postdoctoral Fellow, 1988-1991

Appointments

California Academy of Sciences: Schlinger Chair of Arachnology and Curator (1999-present), Associate Curator (1994-1999), Assistant Curator (1992-1994)
University of California, Berkeley, Adjunct Professor, Insect Biology, Environmental Science, Policy and Management (2004-present)
San Francisco State University, Dept. of Biology, Research Professor (1998-present)
Smithsonian Institution, Research Entomologist (1991-1992)
Natal Museum, Chief Professional Officer, Arachnology Department (1983-1986)

Research Interests

  1. Classification, phylogeny and evolution of spiders, especially as this relates to historical biogeography and to lifestyle (including web) evolution.
  2. Large scale discovery of new taxa, processing of specimens, and the analysis of distribution and diversity patterns.
  3. Historical biogeography, particularly of Gondwana and of the Afromontane region.
  4. Monographs of spiders.
  5. Application of data about richness, diversity and endemism to conservation.

Postdoctoral Associates Advised

Xinping Wang (Schlinger Foundation Postdoctoral Fellow, 2000-2001)
Jeremy Miller (Gaoligongshan Biotic Survey Postdoctoral Fellow, 2207; Schlinger Foundation Postdoctoral Fellow, current)
Diana Silva Dávila (Schlinger Chair of Arachnology Postdoctoral fellow, 2001-2006)
Fernando Alvarez Padilla (Schlinger Chair of Arachnology Postdoctoral fellow, current)

Graduate Students Advised, Current

San Francisco State University
Alma Saucedo (current, major professor)
University of California, Berkeley
Steven Lew (Ph.D., current; committee member)
Natalia Chousou-Polydouri (Ph.D., current; committee member)
Joel Ledford (Ph.D., current; co-advisor and committee member)
Hannah Wood (Ph.D., current; co-advisor and committee member)
The George Washington University
Daniela Andriamalala (Ph.D., current; committee member)
Ligia Benevides (Ph.D., current; committee member)

Graduate Students Advised, Past

San Francisco State University
Joel Ledford (M.Sc., 2001)
Hannah Wood (M.Sc., 2005)
Daniela Andriamalala (M.Sc., 2006)
Misha Leong (M.Sc, 2008), committee member
University of California, Berkeley
Sarah Crews (Ph.D., 2008; committee member)
The George Washington University
Lara Lopardo (Ph.D., 2009; committee member)
University of California at Santa Cruz
Martin G. Ramirez (Ph.D., 1990; committee member)
Rand Afrikaans University, South Africa
Koos de Wet (M.Sc., 1991; committee member)
University of Natal, South Africa
Peter Croeser (M.Sc., 1996; committee member)
University of the Orange Free State, South Africa
Leon Lotz (M.Sc., 1996; committee member)

California Academy of Sciences: Summer Systematics Institute (undergraduate advisees):

Robin Carlson (1996), Joel Ledford (1998), Hillary Guttman (1999) and Nibia Soto-Rolón (2005), Christopher Vo, Vanessa Knutson (2006)

Classes taught:

California Academy of Sciences: Summer Systematics Institute: Lecturer 1996-present; Docent‘s training classes for ‗Life through time‘, ‗Evidence for evolution‘ and ‗Living on the land, Africa.‘
University of California, Berkeley: ESPM 147-2, Field Entomology – spiders (Spring 2006—2009 [current]).
Field Courses taught overseas:
Biology of Arthropoda, lab and field course for Myanmar Department of Forestry, Yangon, Myanmar, September 2003.
Entomological Field Sampling Techniques, Andranomay, Madagascar, December 2000.
Other: Instructor, 1-day field courses on the biology of spiders for East Bay Regional Parks, College of Marin, and Point Reyes Field Seminars, 1980-1983 (all in CA)


Selected Grant Support
:
National Science Foundation (as PI)

DEB-0613775 "PBI: Collaborative Research: The Megadiverse, Microdistributed Spider Family Oonopidae," September 2006 through August 2011 ($319,303)
DEB-0072713: 'Terrestrial Arthropod Inventory of Madagascar' (co P.I. with Brian Fisher) 2000—2003 ($400,917)
DEB 9296271: 'Systematics and Biogeography of Afromontane Spiders,' Sept. 1992-August 1994 ($114,597.00)
BSR 90-20439: 'Systematics and Biogeography of Afromontane Spiders,' April 1991 through June 1992 ($64,074.00.

National Science Foundation (as collaborating Senior Scientist)
EAR-0228699: "Assembling the Tree of Life: Phylogeny of Spiders" (Wheeler, Sierwald, Prendini, Hormiga & Coddington P.I.s), 2003—2008 ($2,700,000)
BSI-0103795: 'Biotic Survey of the Gaoligongshan, a Biodiversity Hotspot in Western Yunnan, China' (N. Jablonski and P. Fritsch, P.I.s), 2002—2007 ($2,400,000)
BIR-9531307: "REU: California Academy of Sciences Summer Systematics Institute" (R. Mooi, T. Gosliner and D. Kavanaugh, P.I.s) with renewals, since 1995.

Other
Lindsay Fund for Expeditions to Madagascar/South Africa (Dec. 2005—March 2006), Myanmar (Sept-Oct. 2003) and Madagascar (December 2008—February 2009)
Lakeside Foundation for International Students:
1999— Balsama Rajemison, Entomology Curator at Parc Tzimbazaza, Madagascar, training in curation techniques for entomological collections for (with B. Fisher).
2000— Daniela Andriamalala, Jean-Jacques Rafanomezantsoa, Balsama Rajemison, Jean Claude Rakotonirina (Madagascar) and Aisha Prather and Madeline van Vyk (South Africa), training in Entomology for four Malagasy and two South Africans (with B. Fisher);
2002— Daniela Andriamalala and Jean-Jacques Rafanomezantsoa (Madagascar): training in Arachnology.
2003— Germania Estevez (Ecuador): training in Arachnology.
2005—Ivalu Avila (Chile): training in Arachnology (with D. Silva).
2005 through 2007— Daniela Andriamalala (Madagascar) M.Sc program at San Francisco State University and preparation and publication of results.
2007—Guo Tang (China): training in Arachnology.
2009—Lina Almeida (Brazil): training in Arachnology.
Central African Research Programs for the Environment (CARPE): Survey of Terrestrial Arthropod Biodiversity on Bioko, Equatorial Guinea (with Robert Drewes), September-October 1998, $17,000.
Research Opportunities Fund: Grant 1233F04A, National Museum of Natural History, to J. Coddington and C. Griswold (for fieldwork in Australia), August 1990.
South African Council for Scientific and Industrial Research support grants: 1985, 1986.

Postdoctoral fellowships

Postgraduate Research Fellowships, Department of Entomology, Smithsonian Institution, July 1988-June 1989 and January 1990-February 1991.
Kalbfleisch Research Fellowships, American Museum of Natural History, January-December 1987 and January-June, 1988.

Student Awards

Exline-Frizzell Fund for Arachnological Research, grant-in-aid, 1985.
Harvy I. Magy Outstanding Graduate Student Award, Department of Entomological Sciences, University of California, Berkeley, 1980.
Theodore Roosevelt Memorial Fellowship, January-February 1978, Department of Entomology, American Museum of Natural History.
Regents Fellowships, University of California, Berkeley, 1976-1977 and 1977-1978.

Other Awards

Paolo Brignoli Award from International Society of Arachnology, 2007.

Field experience:

Conducted fieldwork in Australia, Bolivia, Cameroon, Canada, Chile, China, Costa Rica, Denmark, Guatemala, Madagascar, Mexico, Myanmar, Namibia, New Zealand, Panama, São Tomé e Príncipe, South Africa, Switzerland, Tanzania, Uganda and USA.

Membership in scientific societies:

African Arachnological Society, founding member
American Arachnological Society (Director, 1991-93)
British Arachnological Society
International Society of Arachnology (Vice President, 2001-2004)
Pacific Coast Entomological Society
Society of Systematic Biology
Willi Hennig Society (Fellow since 1989)

Jackson, R. R., and C. E. Griswold. 1979. Nest associates of Phidippus johnsoni (Araneae: Salticidae). The The Journal of Arachnology, 7: 59—67. PDF

Griswold, C. E. 1979. New species of Pellenes from California. The The Journal of Arachnology, 7: 129—138. PDF

Coville, R. E., and C. E. Griswold. 1983. The nesting biology of Trypoxylon xanthandrum (Hymenoptera: Sphecidae) in central America, a hunting wasp specializing on spiders of the family Senoculidae. Journal of the Kansas Entomological Society, 56 (2): 205—216. PDF

Griswold, C. E. 1983. Tapinillus longipes Taczanowski, a web-building lynx spider from the American tropics (Araneae: Oxyopidae). Journal of Natural History, 17: 979—985. PDF

Griswold, C. E. 1984a. The genus Coccorchestes Thorell newly described from Australia (Araneae: Salticidae). Bulletin of the British Arachnological Society, 6: 147—148. PDF

Griswold, C. E. 1984b. A key to the nominate genera of Mygalomorphae (Chelicerata: Araneae) from southern Africa. Newsletter of the spider club of South Africa, 1(5), seperate, 18 p.

Coville, R. E., and C. E. Griswold. 1984. Biology of Trypoxylon superbum (Hymenoptera: Sphecidae), a spider-hunting wasp with extended guarding of the brood by males. Journal of the Kansas Entomological Society, 57: 365—376.
Griswold, curriculum vitae, pg. 7. PDF

Griswold, C. E. 1985a. A revision of the African spiders of the family Microstigmatidae (Araneae: Mygalomorphae). Annals of the Natal Museum, 27: 1—37. PDF

Griswold, C. E. 1985b. Isela okuncana, a new genus and species of kleptoparasitic spider from southern Africa (Araneae: Mysmenidae). Annals of the Natal Museum, 27: 207—217. PDF

Griswold, C. E. and R. E. Coville. 1986. Observations on the prey and nesting biology of spider-hunting wasps of the genus Trypoxylon (Hymenoptera: Sphecidae) in Costa Rica. Proceedings of the IX international Congress of Arachnology (Panama 1983), pp. 113—116. PDF

Griswold, C. E. 1987a. A revision of the jumping spider genus Habronattus F. O. P. Cambridge (Araneae: Salticidae), with phenetic and cladistic analyses. University of California Publications in Entomology, 107: i—ix, 1—344.

Griswold, C. E. 1987b. The African members of the trap-door spider family Migidae (Araneae: Mygalomorphae), I: The genus Moggridgea O. P. Cambridge, 1875. Annals of the Natal Museum, 28: 1—118. PDF

Griswold, C. E. 1987c. The spider genus Symphytognatha Hickman (Araneae: Symphytognathidae) newly described from Africa. Annals of the Natal Museum, 28: 133—136. PDF

Griswold, C. E. 1987d. The African members of the trap-door spider family Migidae (Araneae: Mygalomorphae), 2: The genus Poecilomigas Simon. Annals of the Natal Museum, 28: 475—497. 15. Griswold, C. E. 1987e. A review of the southern African spiders of the family Cyatholipidae Simon, 1894 (Araneae: Araneomorphae). Annals of the Natal Museum, 28: 499—542. PDF

Griswold, C. E. and N. I. Platnick. 1987. On the first African spiders of the family Orsolobidae (Araneae, Dysderoidea). American Museum Novitates, 2892: 1—14. PDF

Griswold, C. E. and T. Meikle Griswold. 1987. Archaeodictyna ulova new species, (Araneae: Dictynidae), a remarkable kleptoparasite of group-living eresid spiders (Stegodyphus spp., Araneae: Eresidae). American Museum Novitates, 2897: 1—11. PDF

Moldenke, A. R., B. L. Fichter, W. P. Stephen and C. E. Griswold. 1987. A key to the arboreal spiders of Douglas-fir and true fir forests of the Pacific Northwest. General Technical Report PNW-GTR-207, Portland. OR: U. S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 48 p.

Griswold, C. E. 1990. A revision and phylogenetic analysis of the spider subfamily Phyxelidinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 196: 1—206.

Griswold, C. E. 1991. Cladistic Biogeography of Afromontane Spiders. Australian Systematic Botany, 4(1): 73—89. PDF

Platnick, N. I., Griswold, C. E., and Coddington, J. 1991. On missing entries in cladistic analysis. Cladistics 7: 337—343. Griswold, curriculum vitae, pg. 8. PDF

Platnick, N. I., J. A. Coddington, R. F. Forster, and C. E. Griswold. 1991. Spinneret evidence and the higher classification of the haplogyne spiders (Araneae, Araneomorphae). American Museum Novitates, 3016: 1—73.

Griswold, C. E. 1991. A revision and phylogenetic analysis of the African spider genus Machadonia Lehtinen (Araneae: Lycosoidea). Entomologica scandinavica, 22: 305—351.

Coddington, J. A., C. E. Griswold, D. Silva-Dávila, E. Peñaranda, and S. Larcher. 1991. Designing and testing sampling protocols to estimate biodiversity in tropical ecosystems. Pp. 44—60 In: Dudley, E. C., (ed.) The Unity of Evolutionary Biology. Proceedings of the Fourth International Congress of Systematic and Evolutionary Biology. Dioscorides Press: Portland, Ore. PDF

Griswold, C. E. 1993. Investigations into the phylogeny of the Lycosoid spiders and their kin (Arachnida, Araneae, Lycosoidea). Smithsonian Contributions to Zoology, 539: 1—39. PDF

Griswold, C. E. 1994. A revision and phylogenetic analysis of the African spider genus Phanotea Simon (Araneae: Lycosoidea). Annales, Sciences Zoologiques, Musée Royal de L’Afrique Centrale, 273: 1—83.

Griswold, C. E. 1997a. The spider family Cyatholipidae in Madagascar (Araneae: Araneoidea). The The Journal of Arachnology 25: 53—83. PDF

Griswold, C. E. 1997b. Scharffia, a remarkable new genus of spiders from East African Mountains (Araneae, Cyatholipidae). The Journal of Arachnology 25: 269—287. PDF

Griswold, C. E., J. Coddington, G. Hormiga, and N. Scharff. 1998. Phylogeny of the orb-web building spiders (Araneae, Orbiculariae: Deinopoidea, Araneoidea). Zoological Journal of the Linnean Society 122: 1—99.

Griswold, C. E. 1998a. The nest and male of the trap-door spider Poecilomigas basilleupi Benoit, 1962 (Araneae, Migidae). The Journal of Arachnology, 26: 142—148. PDF

Griswold, C. E. 1998b. Wanzia fako, a new genus and species of spider from Cameroon (Araneae, Cyatholipidae). Entomologica scandinavica, 29: 121—130. PDF

Griswold, C. E., J. Coddington, N. Platnick, and R. Forster. 1999. Towards a phylogeny of entelegyne spiders (Araneae, Opisthothele, Araneomorphae). The Journal of Arachnology, 27: 53—63. PDF

Griswold, C. E., C.L. Long, and G. Hormiga. 1999. A new spider of the genus Pimoa from Gaoligong Mountains, Yunnan, China (Araneae, Araneoidea, Pimoidae). Acta Botanica Yunnanica 1999, suppl XI: 91—97, plates I—V. PDF

Coville, R. E., C.E. Griswold, and P. L. Coville. 2000. Observations on the Nesting Biology and Behavior of Trypoxylon (Trypargilum) vagulum (Hymenoptera: Sphecidae) in Costa Rica. The Pan Pacific Entomologist, 76(1): 28—48. PDF

Griswold, C. E. 2000 Afromontane spiders in Madagascar (Araneae, Araneomorphae: Cyatholipidae, Phyxelididae, Zorocratidae) pp. 345—354 In: Griswold, curriculum vitae, pg. 9 W. R. Lourenço and S. M. Goodman (Eds.). Diversity and Endemism in Madagascar. Mémoires de la Société de Biogéographie, Paris. PDF

Griswold, C. E. 2001. A monograph of the living world genera and Afrotropical species of cyatholipid spiders (Araneae, Orbiculariae, Cyatholipidae). Memoirs of the California Academy of Sciences, Number 26: 1—251.

Griswold, C. E., and D. Ubick . 2001 Zoropsidae, a spider family newly introduced to the USA (Araneae, Entelegynae, Lycosoidea). The Journal of Arachnology, 28: 111—113. PDF

Griswold, C., and J. Ledford. 2001. A monograph of the migid trap-door spiders of Madagascar, with a phylogeny of world genera (Araneae, Mygalomorphae, Migidae). Occasional Papers of the California Academy of Sciences. Number 151: 1—120. PDF

Xu, X., C. M Yin, and C. E. Griswold. 2002. A New Species of the Spider Genus Macrothele from the Gaoligong Mountains, Yunnan, China (Araneae: Hexathelidae), Pan-pacific Entomologist, 78(2): 116—119. PDF

Griswold, C. E. 2002. A revision of the African spider genus Raecius Simon, 1892 (Araneae, Zorocratidae). Proceedings of the California Academy of Sciences, vol. 53, no. 10: 117—149.

Yin. C. M., C. E. Griswold and H. M. Yan. 2002. A new Ogre-faced spider (Deinopis) from the Gaoligong Mountains, Yunnan, China (Araneae: Deinopidae). The Journal of Arachnology, 30:610—612. PDF

Yin, C. M., C. E. Griswold, Y. H. Bao and X. Xu. 2003. Four Species of the Genus Steatoda from the Gaoligong Mountains, Yunnan, China (Araneae: Theridiidae). Proceedings of the California Academy of Sciences, 54 (6): 133—140. PDF

Griswold, C. E., and H. M. Yan. 2003. On the egg-guarding behavior of a Chinese symphytognathid spider of the genus Patu marples, 1951 (Araneae, Araneoidea, Symphytognathidae). Proceedings of the California Academy of Sciences, 54 (19): 356—360. PDF

Yin, C. M., C. E. Griswold, Y. H. Bao and X. Xu. 2003. A New Species of the Genus Craspedisia from the Gaoligong Mountains, Yunnan, China (Araneae: Theridiidae). Bulletin of the British Arachnological Society, 12(8): 383—384. PDF

Scharff, N., J. Coddington, L. Sørensen, C. E. Griswold, G. Hormiga, and P. de Place Bjørn. 2003. How much is enough? Estimating spider species richness in a northern European deciduous forest. The Journal of Arachnology 31(2): 246—273. PDF

Griswold, C. E. 2003. ―Araneae, Spiders‖, pp. 579—587 In S. Goodman and J. Benstead, eds., The Natural History of Madagascar. University of Chicago Press, Chicago, 1709 p.

Griswold, C. E., M. J. Ramírez, J. Coddington, and N. Platnick. 2005. Atlas of Phylogenetic Data for Entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their Phylogeny. Proceedings of the California Academy of Sciences, 4th Series, vol. 56, Supplement II, pp. 1—324.
Griswold, curriculum vitae, pg. 10.

Tang, G., C. M. Yin, C. E. Griswold and X. J. Peng. 2006. Description of Sinothomisus gen. nov. with a new species from Yunnan Province, China (Araneae, Thomisidae). Zootaxa 1366: 61–68. PDF

Liu, P., H. M. Yan, C. E. Griswold and D. Ubick. 2007. Three new species of the genus Clubiona from China (Araneae: Clubionidae). Zootaxa 1456: 63-68. PDF

Ramírez, M.J., J.A. Coddington, W.P. Maddison, P.E. Midford, L. Prendini, J. Miller, C.E. Griswold, G. Hormiga, P. Sierwald, N. Scharff, S.P. Benjamin, & W.C. Wheeler. 2007. Linking of digital images to phylogenetic data matrices using a morphological ontology. Systematic Biology. 56(2):283—294. PDF

Tang, G., C. M. Yin, X. J. Peng, D. Ubick and C. E. Griswold. 2007. Five crab spiders of the Genus Lysiteles from Yunnan Province, China (Araneae: Thomisidae). Zootaxa 1480: 57-68.

Yin, C. M., C. E. Griswold & X. Xu. 2007. One new species and two new males of the family Araneidae from China (Arachnida: Araneae). Acta arachnol. sin. 16: 1-6.

Wood, H. M., C. E. Griswold, and G. Spicer. 2007. Phylogenetic relationships within an endemic group of Malagasy 'assassin spiders' (Araneae, Archaeidae): ancestor reconstruction, convergent evolution and biogeography. Molecular Phylogenetics and Evolution. 45: 612—619. PDF

Tang, G., X. J. Peng, C. E. Griswold, D. Ubick and C. M. Yin. 2008. Four crab spiders of the family Thomisidae (Araneae, Thomisidae) from Yunnan, China. Acta Zootaxonomica Sinica. 33 (2): 241—247.

Tang, G., C. M. Yin, X. J. Peng, D. Ubick and C. E. Griswold. 2008. The crab spiders of the Genus Lysiteles from Yunnan Province, China (Araneae: Thomisidae). Zootaxa 1742: 1--41. PDF

Wang, Q. B., C. M. Yin, C. Griswold and X. J. Peng. 2009. Three new species of the genus Mallinela (Zodariidae: Araneae) from Yunnan Province, China. Zootaxa 2005: 47–56. PDF

Tang, G., C. M. Yin, X. J. Peng and C. Griswold. 2009. Six crab spiders of the subfamily Stephanopinae from Southeast Asia (Araneae: Thomisidae). The Raffles Bulletin of Zoology, 57(1): 39—50. PDF

Miller JA, Griswold CE, Yin CM (2009a) The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9-195. doi: 10.3897/zookeys.11.160. PDF

Miller JA, Griswold CE, Yin CM (2009b) Appendix C. Locality data (KML format) for all specimens of the spider families Theridiosomatidae, Mysmenidae, Anapidae, and Symphytognathidae collected during an inventory of the Gaoligongshan, Yunnan, China, 1998-2007. DATASET. File format: KML (Keyhole Markup Language) version 2.1 for GoogleEarth. doi: 10.3897/zookeys.11.160-app.C.dt. ZooKeys 11: 9-195. doi: 10.3897/zookeys.11.160

Penev L, Erwin T, Miller J, Chavan V, Moritz T, Griswold C (2009) Publication and dissemination of datasets in taxonomy: ZooKeys working example. ZooKeys 11: 1-8. doi: 10.3897/zookeys.11.210
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Tang, G., C. M. Yin, X. J. Peng. and C. E. Griswold. In press. Description of a new crab spider genus Macroapophysis (Araneae, Thomisidae) from South China. Zootaxa.

Peng, H., C. Griswold, C. M. Yin and X. J. Peng. In press. Two new spider species of the genus Thymoites from Yunnan Province, China (Araneae: Theridiidae). Zootaxa.

Miller, J., C. Griswold and C. Haddad. (in review) Taxonomic Revision of the Spider Family Penestomidae (Araneae, Entelegynae). For Zootaxa.

Almeida-Silva, L., A. Brescovit and C. Griswold. (in review) On the poorly known genus Anuvinda Lehtinen, 1967 (Araneae, Titanoecidae). For Zootaxa.

Miller, J., A. Carmichael, Martín Ramírez, C. Haddad, M. Řezáč, J. Johannesen, J. Král, X. P. Wang , J. Spagna and C. Griswold,. (in review) Phylogenetic Affinities of the Enigmatic Spider Family Penestomidae (NEW RANK; Araneae, Araneoidea, Entelegynae), phylogeny of the Eresidae, and Other Advances in Spider Phylogeny. For Molecular Phylogenetics and Evolution.

Wang, X. P., C. Griswold and D. Ubick. (in review). On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae). for Zootaxa.

Labarque, F., C. Grismado, M. Ramírez, H. M. Yan and C. Griswold. (in review). The Southeast Asian genus Stedocys Ono, 1995 (Araneae: Scytodidae): first descriptions of female genitalia and a new species from China. For Zootaxa. IN PREPARATION

Wood, H., C. Griswold, and A. Carmichael. (in preparation). Molecular phylogeny reveals a single origin for Madagascar‘s endemic lace web spiders (Araneae, Phyxelididae). For Journal of Biogeography.

Ledford, J., and C. Griswold. (in preparation). A redescription of Archoleptoneta schusteri Gertsch (Araneae, Leptonetidae), with a review of the morphology and relationships of the Leptonetidae. For Zootaxa.

Griswold, C. and H. Wood. (in preparation). A monograph and phylogenetic analysis of the lace web spiders (Araneae, Phyxelididae) of Madagascar. For The Zoological Journal of the Linnean Society.

Almeida-Silva, L., A. Brescovit and C. Griswold. (in preparation) On the Asian spider genus Pandava Lehtinen, 1967 (Araneae, Titanoecidae). For Zootaxa.
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Alvarez-Padilla, F, and C. Griswold. (in preparation) A revision of the goblin spider genus Silhouettella Benoit, 1979 from Madagascar (Araneae, Oonopidae). For Bulletin of the American Museum of Natural History.

Bio

Learn about some of Charles Griswold’s work and achievements at the Academy in this brief bio.

Contact

  • Charles Griswold
  • Curator and Schlinger Chair of Arachnology
  • California Academy of Sciences
  • 55 Music Concourse Drive
  • San Francisco, CA 94118
  • 415-379-5312 (direct)