Norm Penny

  • Collections Manager
  • Entomology
  • PhD

Neuroptera and Mecoptera

 

I am interested in the taxonomy, biology, and biogeography of the Mecoptera and Neuropterida.

 

Mecoptera

 

Neuroptera & Megaloptera of Madagascar

 

Philippine Islands Biodiversity Project

B.S., Entomology, Iowa State University (1968)
M.S., Entomology, University of Illinois (1970)
Ph.D., Entomology, University of Kansas (1974)

1975 - 1976: Researcher, U.S. Peace Corps, West Malaysia, and Malaysian Agricultural Research and Development Institute (MARDI) at Rice Research Institute, Bumbong Lima, Province Wellesley, West Malaysia. Duties included research on population dynamics of rice leafhoppers and planthoppers, and their natural predators and parasites.

1976 - 1982: Assistant Researcher, Department of Entomology and Curator, Systematics Research Collections in Entomology, Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brazil. Chairman‚ from 1978 to 1982. Duties included administration of the Department, starting and maintaining the first on-site systematics research collection in Entomology for the upper Amazon region, and graduate level teaching in Portuguese. Research and collecting conducted throughout the Amazon Basin, which included various extended field excursions to remote areas. Consulting studies done at the major Brazilian collections in Belem, Sao Paulo, and Rio de Janeiro. Participation in ecological impact statements for ALCOA, World Bank, and northern Brazil electrical authority.

1982 - Jan. 1984: Associate Researcher and Chief, Department of Entomology and Curator, Systematics Research Collections in Entomology (INPA). Duties the same as previous years.

Feb. - Nov. 1984: Visiting Professor, Department of Entomology, Iowa State University, Ames, Iowa. Full time research on corn insect ecology. Duties included assisting Dr. William B. Showers with research on ecology, bionomics, and migration of corn insects, principally Black Cutworm, Armyworm, and European Corn Borer. Studies included monitoring seasonal agricultural insect population dynamics for potential allergic reactions.

Dec. 1984 - Jan. 1991. Collections Manager, Department of Entomology, California Academy of Sciences. Duties include computerization of collection holdings, loans, and accessions, using Microsoft’s ACCESS relational database software. Other duties include identification of insects for the public, supervision of departmental technicians, presenting public talks, assisting visitors and guests, assisting in writing departmental grant proposals and reports, and upgrading the curation of the Entomology Collection.

Feb. 1991 - present. Senior Collections Manager, Department of Entomology, California Academy of Sciences. Duties include same as above for Collections Manager.

TEACHING EXPERIENCE:

1973 - 1974. Graduate Teaching Assistant, General Entomology, University of Kansas. Duties included supervising laboratories and occasionally giving short lectures on general Entomology topics.

1977, 1978, 1980, 1982. Professor, Insect Taxonomy (INPA - Manaus. Topics included taxonomy of Brazilian insects with introduction to the principles of systematics and nomenclature; instruction in Portuguese.

1979, 1981. Professor, Insect Evolution (INPA - Manaus). Topics included evolution of extinct and extant insectan groups and their morphological structures; instruction in Portuguese.

1990. Instructor, Insects of Medical Importance. A CAS adult short course taught one night a week for one month (Jan. 9, 16, 23, 30). Each class was for two hours and dealt with current public health issues such as Lyme Disease, Plague, Malaria, etc.

1991. Instructor, Insects of the Tropics. A CAS adult short course taught Oct. 21, 1991. Colorful tropical insects were shown in slides and preserved specimens.

1992. Instructor, "Species Lost". A team taught CAS adult short course taught on two evenings for 2 hours each (Feb. 6, 13). Half hour presentations on various groups of endangered or extinct species.

1995. Instructor, "Taxonomy of Neuropteroid Insects". A one credit course sponsored through San Francisco State University on the detailed taxonomy of Neuroptera. Course given over five Saturdays from April 22 to May 20, 1995. Total 10 hours lecture, 30 hours lab.

l966. Sparks, A.N., Brindley, T.A. and Penny, N.D. Laboratory and field studies of F1 progenies from reciprocal matings of the European Corn Borer. Journal of Economic Entomology, 59:915-921.

 

l969. A new species of “Bittacus" (Mecoptera: Bittacidae) from India. Oriental Insects, 3:161-164.

 

l975. Webb, D.W., Penny, N.D. and Marlin, J.C. The Mecoptera, or scorpion-flies, of Illinois. Bulletin of the Illinois Natural History Survey, 31:251-316, 187 figs.

 

1975. Evolution of the extant Mecoptera. Journal of the Kansas Entomological Society, 48:331-350, 7 figs.

 

1977. A systematic study of the family Boreidae (Mecoptera). University of Kansas Science Bulletin, 51:141-217. 

 

1977. Problems and solutions to insect identification and literature retrieval in developing countries (abstract). Association of Systematic Collections
Newsletter, 5(3):29.

 

1977. Two new species of Bittacidae (Mecoptera) from the Amazon Basin. Acta Amazonica, 7(3):423-427. 

 

1978. Lista de Megaloptera, Neuroptera e Raphidioptera do Mexico, America Central, Ilhas Caraibas e America do Sul. Acta Amazonica, (supl.), 7(4):1-61.

 

1978. Webb, D.W. and Penny, N.D. “Neopanorpa byersi" (Mecoptera: Panorpidae) a new species from Thailand. Pacific Insects, 20:63-66.

 

1978. Penny, N.D. and Avery, M.L. Collections of scorpionflies (Mecoptera) of West Malaysia. Malay Nature Society Journal, 30:555-559. 

 

1978. Avery, M.L. and Penny, N.D. Analysis of pellets from Blue-tailed and Blue- throated Bee-eaters in ProvinceWellesley, Malaysia. Malay Nature Society Journal, 30: 

 

1978. Penny, N.D., Arias, J.R. and Schubart, H.O.R. Tendencias populacionais da fauna de Coleopteros do solo sob floresta da terra firma na Amazonia. Acta Amazonica, 8(2):259-266.

 

1978. Ratcliffe, B.C. and Penny, N.D. As colecoes de Entomologia Sistematica do Instituto Nacional de Pesquisas da Amazonia. Ciencia e Cultura, 30(5):627-628.

 

1978. Ratcliffe, B.C. and Penny, N.D. Catalogo dos tipos de invertebrados depositados na Colecoes Entomologicas do INPA-Manaus. Acta Amazonica, 8(4):689-698.

 

1979. Penny, N.D. and Byers, G.W. A check-list of the Mecoptera of the World. Acta Amazonica, 9(2):365-388.

 

1979. Penny, N.D. and Byers, G.W. Chave para a identificacao das familias e generos da ordem Mecoptera do Mexico, America Central e America do Sul. Acta Amazonica, 9(2):362-364.

 

1980. A revision of American Bennini (Hemiptera: Fulgoroidea: Cixiidae). Acta Amazonica, 10(1):97-102.

 

1980. Arias, J.R. and Penny, N.D. A entomofauna da Bacia Amazonica. Acta Amazonica, (supl.), 9(4):103-107.

 

1981. Penny, N.D. and Arias, J.R. Two new species of “Bakerius" from the Amazon Basin (Homoptera: Aleyrodidae). Acta Amazonica, 10(3):691-695.

 

1981. Penny, N.D. and Arias, J.R. A new species of “Ceraleurodius" from the Amazon Basin (Homoptera: Aleyrodidae). Acta Amazonica, 10(4):903-905.

 

1981. Penny, N.D. Neuroptera, In, Hurlburt, S.B., Rodriguez, G. and dos Santos, N.D. Aquatic Biota of Tropical South America. Part 1 Arthropoda. San Diego State University Press, San Diego, CA, U.S.A. 323 pp.

 

1981. Neuroptera of the Amazon Basin. Part 1 Sisyridae. Acta Amazonica, 11(1):157-169.

 

1981. Neuroptera of the Amazon Basin. Part 2 Dilaridae. Acta Amazonica, 11(2):383-390.

 

1981. Review of the generic level classification of New World Ascalaphidae (Neuroptera). Acta Amazonica, 11(2):391-406.

 

1981. Neuroptera of the Amazon Basin. Part 3 Ascalaphidae. Acta Amazonica, 11(3):605-651.

 

1981. Penny, N.D. and Arias, J.R. Observations on the egg and larva of “Nannobittacus elegans" Esben-Petersen, 1927 (Mecoptera: Bittacidae). Journal of the Kansas Entomological Society, 54(3):465-468.

 

1982. Neuroptera, pp. 280-282, In, Hurlburt, S.B. and Villalobos-Figueroa, A., Aquatic Biota of Mexico, Central America, and the West Indies. San Diego State University Press, San Diego, CA, U.S.A.

 

1982. Neuroptera of the Amazon Basin. Part 4 Sialidae. Acta Amazonica, 11(4):843-846.

 

1982. Review of the generic level classification of New World Mantispidae (Neuroptera). Acta Amaznica, 12(1):209-223.

 

1982. Penny, N.D. and Arias, J.R. Insects of an Amazon Forest. Columbia University Press, New York. 270 pp., 127 figs., 93 graphs, and 18 tables.

 

1982. Penny, N.D. and Flint, O.S., Jr. Revision of the genus “Chloronia" (Megaloptera: Corydalidae). Smithsonian Contributions to Zoology, 348:1-27.

 

1982. Meinander, M. and Penny, N.D. Neuroptera of the Amazon Basin. Part 5 Coniopterygidae. Acta Amazonica, 12(1):185-208.

 

1982. Neuroptera of the Amazon Basin. Part 6 Mantispidae. Acta Amazonica, 12(2):416-453.

 

1982. Notes on Amazonian Mecoptera with descriptions of two new species. Memorias do Instituto Oswaldo Cruz, 77(3):263-274.

 

1983. Neuroptera of the Amazon Basin. Part 7 Corydalidae. Acta Amazonica, 1294):825-837.

 

1983. Monserrat, V.J. and Penny, N.D. Sobre las especies americanas del genero “Notiobiella" Banks, 1909, II (Neuroptera, Planipennia, Hemerobiidae). Nouv. Rev. Ent., 13:125-132.

 

1983. Penny, N.D. and Rafael, J.A. Two new species of Sisyridae (Neuroptera) from the Amazon Basin. Neuroptera International, 2(2):53-58.

 

1984. Penny, N.D. and Sturm, H. A new flightless brown lacewing (Neuroptera: Hemerobiidae) from Colombia. Spixiana, 7(1):19-22.

 

1984. Recent progress in the Taxonomy of South American Neuroptera, p. 75-77, In, Gepp., J., Aspoeck, H., and Hoelzel, H., Progress in World's Neuropterology. Graz, Austria. 265 pp.

 

1984. Penny, N.D. and Costa, C.A. Mantispideos do Brasil (Neuroptera: Mantispidae). Acta Amazonica, 13(3/4):601-688.

 

1984. Neuroptera of the Amazon Basin. Part 8 Berothidae. Acta Amazonica, 13(3/4):689-696.

 

1984. Neuroptera of the Amazon Basin. Part 9 Albardiinae. Acta Amazonica, 13(3/4):697-700.

 

1984. Penny, N.D. and Arias, J.R. A new species of “Issikiella" from the Amazon Basin (Mecoptera: Bittacidae). Acta Amazonica, 13(3/4):701-704.

 

1985. Rafael, J.A., Penny, N.D. and Harada, A.Y. Relacao dos tipos de invertebrados depositados na Colecao de Entomologia Sistematica do INPA- Manaus, ate dezembro de 1982. Acta Amazonica, 13(5-6):911-926.

 

1985. Penny, N.D. and Adis, J. A blind Mecoptera larva from Reserva Ducke. Acta Amazonica, 13(5-6):875-877.

 

1985. Carico, J.E., Adis, J. and Penny, N.D. A new species of “Trechalea" (Pisauridae, Araneae) from Central Amazonian inundation forests and notes on its natural history and ecology. Bulletin of the British Arachnological Society, 6(7):289-294.

 

1985. Penny, N.D. and Monserrat, V.J. Neuroptera of the Amazon Basin. Part 10 Hemerobiidae. Acta Amazonica, 13(5-6):879-909.

 

1985. Penny, N.D. and Ratcliffe, B.C. Entomological Collections and Human Resources in Brazil. Assoc. of Systematic Collections Newsletter, 13(3): 21-24.

 

1986. Penny, N.D. and Arias, J.R. Four new species of wing rolling “Neodawnaria" (Homoptera: Derbidae) from the Amazon Basin. Acta Amazonica, 14(3/4):459- 471.

 

1986. Adams, P.A. and Penny, N.D. Faunal Relationships of Amazon Chrysopini, pp. 119-124, In, Gepp, J., Aspoeck, H and Hoelzel, H. Recent Research in Neuropterology. Graz, Austria. 176 pp.

 

[1987] 1985. Adis, J., Junk, W.J. and Penny, N.D. Material zoologico depositado pelo Instituto Max-Planck de Limnologia nas Colecoes Sistematicas de Entomologia do Instituto Nacional de Pesquisas da Amazonia (INPA) Manaus, Brasil. Acta Amazonica, 15(3-4):481-504.

 

[1987] 1985. Adams, P.A. and Penny, N.D. Neuroptera of the Amazon Basin.Part 11a. Introduction and Chrysopini. Acta Amazonica, 15:413-479.

1988. Computer breakdown at CAS. Insect Collection News, 1(1):9.

 

[1988] 1987. O'Brien, L.B., Penny, N.D., and Arias, J.R. Lophopidae of the Amazon Basin with keys to New World Genera and species (Homoptera: Fulgoroidea). Acta Amazonica, 16/17:617-626.

 

1988 Wynn, S.R., Swanson, M.C., Reed, C.E., Penny, N.D., Showers, W.B., Smith, J.M. Immunochemical quantitation, size distribution, and cross-reactivity of Lepidoptera (moth) Aeroallergens in Southeastern Minnesota. Journal of Allergy and Clinical Immunology, 82(1):47-54.

 

[1989] 1988 Hogue, C.L. and Penny, N.D. Aggregations of Amazonian Owlflies (Neuroptera: Ascalaphidae:Cordulecerus). Acta Amazonica, 18(1-2):359-362.

 

1989. Penny, N.D. Effects of the recent earthquake on the CAS Entomological Collection. Insect Collection News, 2(2):4-5.

 

1991. Oswald, J.D. and Penny, N.D. Generic Names of the Neuroptera, Megaloptera and Raphidioptera of the World. Occasional Papers of the California Academy of Sciences. No. 147, 94 pp.

 

1992. Henry, C.S., Penny, N.D., and Adams, P.A. The neuropteroid orders of Central America (Neuroptera and Megaloptera), pp. 432-458, In, Quintero, D. and Aiello, A., editors, Insects of Panama and Mesoamerica, selected studies. Oxford University Press, Oxford, England. 692 pp.

 

1992. Adams, P.A. and Penny, N.D. Review of the South American genera of Nothochrysinae (Insecta: Neuroptera: Chrysopidae), In, Canard, M., Aspoeck, H., and Mansell, M.W., Current Research in Neuropterology. 414 pp.

 

1992. Adams, P.A. & N.D. Penny. New genera of Nothochrysinae from South America (Neuroptera: Chrysopidae). Pan-Pacif. Entomol., 68(3):216-221. 

 

1993. Penny, N.D. The phylogenetic position of Chloroniella peringueyi Esben- Petersen and its zoogeographic significance. Entomological News, 104:17-30.

 

1993. Penny, N.D. [Book Review] Sarapiqui Chronicle by Allen M. Young. American Entomologist. 38(2):

 

1994. Penny, N.D. A new species of Nallachius from Costa Rica. Pan-Pacif. Entomol. 70(4):309-312.

 

1995. Penny, N.D. Portraits in Lace. Pacific Discovery. 48(4):28-31.

 

1995. Penny, N.D. Specimen Level Databases. Insect Collection News 10:7-9.

 

1996. Penny, N.D. and Vincent F. Lee. Types of Mecoptera, Megaloptera, Raphidioptera, and Neuroptera in the Collection of the California Academy of Sciences. Transactions of the California Academy of Sciences 49(4):127-149.

 

1996. Penny, N.D. A remarkable new genus and species of Ithonidae from Honduras (Neuroptera). Journal of the Kansas Entomological Society 69(1): 81-86.

 

1996. Penny, N.D. A Revision of Parachrysopiella (Neuroptera: Chrysopidae). Journal of the Kansas Entomological Society 69(3):222-233.

 

1997. Penny, N.D. Four new species of Costa Rican Ceraeochrysa. Pan-Pacific Entomologist 73(2):61-69.

 

1997. Penny, N.D., P.A. Adams, and L.A. Stange. Species Catalog of the Neuroptera, Megaloptera, and Raphidioptera of America north of Mexico. Proceedings of the California Academy of Sciences 50(3):39-114.

 

1998. Penny, N.D. New Chrysopinae from Costa Rica (Neuroptera: Chrysopidae). Journal of Neuropterology 1:55-78.

 

1998. Penny, N.D. The Neuroptera of Costa Rica – an Overview. Proceedings of the VIth International Symposium of Neuropterology. Acta Zoologica Fennica 209:211-214.

 

1998. Penny, N.D. [Review] Zoological Catalogue of Australia. Neuroptera, Strepsiptera, Mecoptera, Siphonaptera, vol. 28. Wells, A. [ed.] CSIRO Publishing, Victoria, Australia, 232 pp.

 

1999. Zack, R.S., N.D. Penny, J.B. Johnson, and D.L. Strenge. Raphidioptera and Neuroptera from the Hanford Site of Southcentral Washington State. Pan-Pacific Entomologist 74(4):203-209.

 

1999. Madachauliodes ranomafana, a new Madagascan species of Chauliodinae (Megaloptera: Corydalidae), with a key to the world genera of Chauliodinae. Entomological News 110(4):193-200.

 

1999. Allen, R.E., N.D. Penny and J.D. Oswald. Obituary: Phillip A. Adams. Journal of Neuropterology 2:5-86.

 

2000. Penny, N.D., C.A. Tauber and T. De Leon. A new species of Chrysopa from western North America with a key to North American species (Neuroptera: Chrysopidae). Annals of the Entomological Society of America 93(4):776-784.

 

2000. Tauber, C.A., T. De Leon, N.D. Penny, and M.J. Tauber. The genus Ceraeochrysa (Neuroptera: Chrysopidae) of America north of Mexico. Annals of the Entomological Society of America 93(6):1195-1221. 16 figures. 2 tables.

 

2001. Penny, N.D. New Species of Chrysopinae (Neuroptera: Chrysopidae) from Costa Rica with selected taxonomic notes and a neotype designation. Entomological News 112(1):1-14.

 

2001. S. de Freitas and N.D. Penny. Two New Genera of Neotropical Chrysopini (Neuroptera: Chrysopidae). Journal of the Kansas Entomological Society 73(3):164-170.

 

2001. Penny, N.D. and S. de Freitas. Sexual Dimorphism in Ceraeochrysa cubana (Neuroptera: Chrysopidae). Journal of the Kansas Entomological Society 74(1):38-39.

 

2001. S. de Freitas and N.D. Penny. The Green Lacewings (Neuroptera: Chrysopidae) of Brazilian Agro-Ecosystems. Proceedings of the California Academy of Sciences 52(19):245-395.

 

2002. Contreras-Ramos, A., J.D. Oswald and N.D. Penny. Chapter 33. Neuroptera (Neuropterida) In, J.L. Bousquets and J.J. Morrone, eds., Biodiversidad, Taxonomía y Biogeografía de Artrópodos de México: Hacia una sítesis de su conocimiento. Volume 3. Universidad Nacional Autónoma de México, Ciudad Universitaria, México. 690 pp.

 

2002. Penny, N.D. et al. A Guide to the Lacewings (Neuroptera) of Costa Rica. Proceedings of the California Academy of Sciences 53(12):161-457.

 

2004. Penny, N.D. Chapter - Megaloptera. Pp. 661-663 In Goodman, S. and J. Benstead, A Natural History of Madagascar. University of Chicago Press.

 

2004. Penny, N.D. Chapter – Neuroptera. Pp. 663-669 In Goodman, S. and J. Benstead, A Natural History of Madagascar. University of Chicago Press.

 

2005. Penny, N.D. The California Academy of Sciences Gulf of Guinea Expedition (2001). IV. The Neuroptera of São Tomé and Principé Islands. Proceedings of the California Academy of Sciences 56(22):285-293. 4 figs., 1 table.

 

2006. Penny, N.D. Two New Species of Meleoma (Neuroptera: Chrysopidae) from Western North America, with a Revised Key to Species. Annals of the Entomological Society of America 99(1):58-63.

 

2006. Penny, N.D. A Review of our Knowledge of California Mecoptera. Proceedings of the California Academy of Sciences 57:365-372.

 

2006. Couri, M. and N.D. Penny. On Palpibracus Rondani (Diptera: Muscidae) with Descriptions of New Species. Proceedings of the California Academy of Sciences 57 (15): 451-471.

 

2006. Couri, M.S., A.C. Pont, and N.D. Penny. Muscidae (Diptera) from Madagascar: Identification Keys, Descriptions of New Species, and New Records. Proceedings of the California Academy of Sciences 57(29):799-923, 1 plate, 120 Figs.

 

2007. Penny, N.D., J.R. Arias, and J.S. Johnson. Seasonal Emergence of Neuroptera in Fairfax, County, Virginia. Proceedings of the California Academy of Sciences 58(2):7-19. 32 figs.

 

2007. Penny, N.D. and S.L. Winterton. Rediscovery of the Unusual Genus Ormiscocerus (Neuroptera: Berothidae: Cyrenoberothinae). Proceedings of the California Academy of Sciences 58(1):1-6. 6 figs.

 

2007. Couri, M.S. and N.D. Penny. Redescription and Additional Records of Arthurella Albuquerque (Diptera: Muscidae:Cyrtoneurininae). Proceedings of the California Academy of Sciences 58(6):99-103. 8 figs.

 

2008. Fisher, B.L. and N.D. Penny. Les arthropodes. In, Paysages naturels et biodiversité de Madagascar, S.M. Goodman editor. Publications scientifiques du Museum. Paris, France. 694 pp.

 

(accepted). Penny, N.D. and S. de Freitas. Neuropteros de Reserva Ducke. In, Os Insetos de Reserva Ducke.

 

(accepted). Freitas, S. and N.D. Penny. Neuroptera. In, Os Insetos do Brasil.

 

(accepted). Penny, N.D. Mecoptera. In, J. Adis and J.R. Arias editors, Aquatic Biota of Latin America. Pensoft.

 

(accepted). Penny, N.D. Neuroptera. In, J. Adis and J.R. Arias editors, Aquatic Biota of Latin America. Pensoft.

 

(submitted). Freitas, S. and N.D. Penny. A revision of the New World genus Ceraeochrysa (Neuroptera: Chrysopidae). Proceedings of the California Academy of Sciences.

Contact

  • Norm Penny
  • Collections Manager
  • California Academy of Sciences
  • 55 Music Concourse Drive
  • San Francisco, CA 94118
  • 415-379-5320 (direct)