Voucher Specimens Used In:

Clinal Variation in the Halesia carolina Complex (Styracaceae).

Systematic Botany Volume 25.
[Accepted August 1999]

PETER W.FRITSCH and SHANNON D. LUCAS1

Department of Botany, California Academy of Sciences, San Francisco, CA 94118

1 Present Address: Wetlands Research Associates, Inc., 2169-G East Francisco Blvd., San Rafael, CA 94901


The following is voucher data for the publication Fritsch, Peter W. and Shannon D. Lucas, 2000, "Clinal Variation in the Halesia carolina Complex (Styracaceae)," Systematic Botany volume 25. Specimens examined include those used for the morphological analysis and construction of the distribution map in Fig. 1. Specimens used for the morphological analysis are indicated by "fl" or "fr" following the herbarium acronym; fl = flowering specimens; fr = fruiting specimens. Only one specimen is listed for each county except in cases where more than one collection from a county was used in the morphological analysis. Specimens from the following herbaria were used: A, F, FSU, GA, GH, LL, LSU, MO, NCU, NY, TENN, TEX, US.

Alabama
 
Baldwin Co.: Fairhope, 29 Mar 1924, Jack 3024 (GA [fl]), 2 mi N of US 31 at Spanish Fort above Buzbees Fishing Camp, 27 Mar 1971, Kral 41,853 (F, GA [fl]).
 
Barbour Co.: Eufaula National Wildlife Refuge, 17 Mar 1982, Folsom & Brown 10,292 (TEX); Bibb Co., ca. 6 mi N of Centreville at Little Schultz Creek, 12 Apr 1992, Hill 22,945 (NY [fl], TEX).
 
Blount Co.: 2.3 mi W of Cleveland Community along Locust Fork of Black Warrior River, 5 Sep 1966, Clark 8526 (NCU).
 
Butler Co.: 2.9 mi W of McKenzie, 21 Jun 1967, Clark 14,513 (NCU).
 
Calhoun Co.: 1.5 mi N of Hicks community along Choccolocco Creek, 6 Sep 1966, Clark 8531 (NCU).
 
Cherokee Co.: 4.5 mi from junction of DeKalb Co. rte. 81 with AL rte. 35 along slope of Little River Canyon nr. stream, 11 Aug 1957, Sherman 67 (TENN [fr])
 
Chilton Co.: ca. 10 mi NNE of Verbena, 6 Jun 1972, Kral 47303 (MO).
 
Clay Co.: along Cheaha Creek and Lake Chinnabee in Talladega National Forest, 10 Apr 1981, Whetstone & Bollinger 10,506 (NCU).
 
Cleburne Co.: 1.5 mi S of junction of Skyway and co. 55, S of Piedmont, 22 Jul 1968, Kral 31,836 (FSU).
 
Colbert Co.: ravine above Tennessee River on TVA Wilson Dam Reservation, 31 Aug 1966, Clark & Landers 7885 (NCU [fr]).
 
Conecuh Co.: in the Red Hills area 5 mi SE of Evergreen at Old Beulah Cemetery along hwy. 29, 21 Mar 1985, Diamond 88 (GA [fl]).
 
Covington Co.: SW of Andalusia in Blue Springs Wildlife Management Area in Conecuh National Forest, 17 Jul 1970, Kral 40,078 (GA [fr]).
 
Crenshaw Co.: along Patsaliga River, 8.7 mi W of Brantley, 22 Jun 1967, Clark 14846 (NCU).
 
Cullman Co.: Eight Miles Creek, 21 Jun 1897, Eggert s.n. (MO, NY, US).
 
Dale Co.: 3 mi N of Pinckard nr. Choctawhatchie River, 25 Mar 1967, McDaniel 8664 (FSU, TENN).
 
Dallas Co.: nr. Selma, 11 Apr 1912, Harbison [10]822 (NCU, TENN [fl]).
 
De Kalb Co.: ravine below Sand Mountain off AL 68, 11 Apr 1969, Kral 34,169 (MO, TENN).
 
Escambia Co.: W side Brewton on US 29 in Conecuh River bottoms, 30 Mar 1968, Kral 29,995 (GA).
 
Etowah Co.: 1.7 mi SE of Gallant Community at natural bridge over Little Canoe Creek, 7 Sep 1966, Clark & Heard 8573 (NCU).
 
Geneva Co.: 4 mi NW of Hartford along the Choctawhatchie River, 25 Mar 1967, McDaniel 8650 (FSU, GA [fl], TENN).
 
Greene Co.: 6.8 mi SSE of Aliceville along Sipsey River, 9 May 1969, McDaniel 11,641 (FSU, GA).
 
Hale Co.: 9 Apr 1966, Williams 203 (NCU [fl]).
 
Henry Co.: E of Co. Rd. 7 along East Choctawhatchee River, 28 Jul 1966, Clark 6226 (NCU).
 
Jackson Co.: Pisgah, 12 Sep 1899, Harbison 1036 (NCU).
 
Jefferson Co.: 0.5 mi W of Village Creek along Village Creek, 24 Jun 1949, Harvill 5075 (GA, TEX [fr]).
 
Lee Co.: Auburn, 5 Apr 1931, Ashe s.n. (NCU [fl]), 4 Apr 1896, Earle s.n. (NY [fl]), 24 Mar 1897, Earle & Baker s.n. (NY, US [fl]), 27 Mar 1897, Earle & Baker s.n. (MO, NY [fl]), 11 Apr 1900, Earle & Earle 55 (NY, US [fl]), Lake Wilmore, 27 Jul 1958, Williams s.n. (FSU [fr]).
 
Macon Co.: vic. of Sandy Springs, 11 Apr 1966, Koelling s.n. (NCU).
 
Marengo Co.: 1 mi N of Linden on US 43, 1 Apr 1968, Kral 30,021 (FSU, NCU [fl], TENN).
 
Marshall Co.: W of Nixon off AL 79 above Big Spring Creek, 11 Apr 1969, Kral 34,207 (NCU, TENN).
 
Montgomery Co.: Teasby's Mill, 15 May 1914, Howell 829 (US).
 
Morgan Co.: nr. Decatur along Tennessee River, Apr 1864, Stewart s.n. (F [fl]).
 
Perry Co.: 14.7 mi S of Bibb Co. line on AL 219, 1 Apr 1972, Kral 45459 (MO).
 
Pike Co.: 5 mi ESE of Troy nr. Big Pocosin, 28 Sep 1966, McDaniel 7829 (FSU [fr]).
 
Randolph Co.: SE of Wadley on AL 77, 31 Mar 1967, Kral 28,037 (FSU, TENN).
 
Saint Clair Co.: E of Lake Sumtanga, 12 Apr 1963, Bostick 282—4 (NCU [fl]).
 
Shelby Co.: Oak Mountain State Park, 29 Mar 1973, Kral 49,438 (MO [fl]), 1875, Everts s.n. (F [fl]).
 
Talladega Co.: nr. Clay Co. line along Salt Creek dam site #18, 20 Jun 1966, Clark & Heard 2913A (NCU [fr]), 20 Jun 1966, Clark & Heard 2913B (NCU [fr]).
 
Tallapoosa Co.: S of Dadeville, 11 Apr 1965, Koelling 2183 (NCU, TENN).
 
Tuscaloosa Co.: Warrior River, 10 Apr 1926, collector not indicated, 283 (F [fl]), forested valley side of Black Warrior River, 29 Mar 1977, Sisk 92 (NCU [fl]), Apr 1892, Ward s.n. (NCU, US [fl]), right bank of Warrior River 5 mi above Tuscaloosa, 11 Apr 1892, Ward s.n. (US [fl]), NE part of co. along Hurricane Creek, 3 Apr 1963, Williams 35 (NCU [fl]).
 
Walker Co.: between Summiton and Jasper above Warrior River nr. where crossed by US 78, 13 Apr 1969, Kral 34,274 (GA [fl], MO, US).
 
Arkansas
 
Cleburne Co.: Heber Springs, 31 Oct 1914, Palmer 6978 (MO [fr], US).
 
Franklin Co.: 4 May 1958, Clewell 165B (FSU [fl]).
 
Garland Co.: nr. Ouachita River at Hot Springs, 22 Apr 1924, Palmer 24,486 (MO [fl]).
 
Johnson Co.: Hagarville, 11 Aug 1938, Demaree 19,808 (NY [fr]), N of Hagarville along Peters Branch, 16 Apr 1967, Tucker 3990 (NCU [fl]).
 
Polk Co.: N side of Rich Mountain nr. Rich Mountain Station, 2 Oct 1923, Palmer 23,968 (MO [fr], US), Rich Mountain ca. 0.5 mi above US hwy. 270, 11 Apr 1948, Robbins 2902 (NY [fl]).
 
Pope Co.: Dover, 11 Aug 1939, Demaree 19,844 (MO [fr], NY), 15 mi W of Mena on AR 88 in Wilhelmina State Park, 1 Oct 1988, Hess & van der Linden 6412 (GA [fr], NCU).
 
Saline Co.: Deep shaded woods bottom, 21 Oct 1965, Demaree 53,155 (NCU).
 
Florida
 
Bay Co.: 1.2 mi W of Bennett along Econfina Creek, 4 Aug 1967, McDaniel 9619 (FSU [fr]).
 
Calhoun Co.: E of Clarkesville on hwy. 20 bordering Chipola River, 29 Apr 1981, Anderson 5195 (FSU).
 
Clay Co.: Hibernia/Magnolia, Mar—Apr 1858, Canby s.n. (NY [fl]), Hibernia, Mar 1869, Canby s.n. (NY [fl], US), borders of swamp along St. John’s River, 12 Jun 1884, Lewis s.n. (US [fr]), nr. Magnolia, Sep 1884, Lewis s.n. (US [fr]), Governor’s Creek, 28 Feb 1883, Smith s.n. (F [fr], US [fl]), nr. Magnolia, 10 May 1884, Smith s.n. (NY [fr]), swamp nr. Magnolia, 27 Mar 1886, Smith s.n. (F, NY, US [fl]).
 
Columbia Co.: 2 Nov 1941, collector not indicated, s.n. (NY).
 
Dixie Co.: along Suwannee River at wayside park W of the River beyond Fanning Springs, 27 May 1981, Godfrey 78,770 (FSU).
 
Escambia Co.: NE of Cantonment along the Escambia River, 1.5 mi N of rte. 184, 31 Mar 1984, Anderson 6890 (FSU [fl]), NE of Atmore along Escambia Creek at I—65, 27 Jul 1969, Kral 35,835 (FSU [fr]).
 
Gadsden Co.: banks of Apalachicola River nr. Chattahoochee, 12 Mar 1897, Biltmore Herbarium No. 520b (NY, US [fl]), 2 mi E of Havana, 1 Apr 1958, Godfrey 56,234 (FSU, GA, LL, MO, NCU [fl]), NY, TENN, TEX, US), E of Quincy along the Little River by US rte. 90, 27 Mar 1981, Godfrey 78,473 (FSU [fl]), River Junction, 27 Mar 1914, Harbison 11,410 (NCU [fl], NY), 10—11 Aug 1895, Nash 2373 (F [fr], MO, NCU, NY, US).
 
Gilchrist Co.: along the Suwannee River at Fanning Springs, 25 Mar 1958, Godfrey & Kral 56,305 (FSU [fl]).
 
Gulf Co.: E of Wewahitchka along the Apalachicola River, 11 Mar 1982, Godfrey 79,373 (FSU [fl]).
 
Hamilton Co.: 2 mi E of Jennings on FL hwy. 150 along W side of Alapaha River, 27 Oct 1976, Zeigler 95 (GA [fr], NCU).
 
Holmes Co.: 4.5 mi SW of Millers Crossroads, 4 Apr 1958, Godfrey 56,401 (FSU).
 
Jackson Co.: banks of Chipola River between Marianna and Florida Caverns State Park, 20 Jul 1966, Chester & Chester 1350 (TENN [fr]), along the Chipola River between Marianna and Marianna Caverns State Park, 29 Mar 1958, Godfrey 56,318 (NCU [fl], NY), E of Clarkesville nr. the Chipola River, 3 Apr 1958, Godfrey 56,337 (FSU [fl]), N of Marianna toward the N end of Bumpnose Rd., 0.5 mi N of Waddell’s Mill Creek, 26 Mar 1984, Godfrey & Gholson 81,167 (FSU [fl]), Marianna, 31 Mar 1940, Henry 2001 (GA [fl]), 0.5 mi NE of hwy. 90 bridge over Chipola River in Marianna, 2 Apr 1979, Whitten et al. 732 (TENN [fl]), S of Marianna 100 yd from Chipola River at Dry Creek, 4 Jun 1957, Williams s.n. (FSU [fr]).
 
Lafayette Co.: along Suwannee River, 13 Sep 1918, Harbison s.n. (NCU [fr]), Old Town, 31 Mar 1920, Harbison 15,622 (NCU [fl]).
 
Leon Co.: along Ochlockonee River down river of Rock Bluff by Forest Rd. 390 N of rte. 375, 22 Mar 1988, Anderson 11,119 (FSU [fl]), nr. hwy. 12 nr. small creek E of Havanna, 2 Apr 1966, Chester & Chester 1036 (TENN [fl]), W of Havanna along FL hwy. 12 nr. small stream, 20 Jul 1966, Chester & Chester 1293 (TENN [fr]), along Ochlockonee River at Larkin’s Bluff below Talquin Dam, 15 Mar 1956, Godfrey 54,468 (FSU, TENN [fl]).
 
Liberty Co.: Torreya State Park, 4 Apr 1972, Elliot 34 (FSU), nr. White Springs, Jun 1963, Godfrey & Houk 62,848a (FSU [fr]).
 
Okaloosa Co.: N of ERR 456 on Eglin Air Force Base, 23 Jul 1992, Anderson 13,750 (FSU [fr]), Rocky Run, 5 Aug 1928, Ashe s.n. (NCU [fr]).
 
Saint John's Co.: Pellicier’s Creek, 2 Mar 1882, Smith s.n. (F [fl], NY, US).
 
Santa Rosa Co.: along Escambia River, W of Jay, 7 Mar 1957, Godfrey & Harrison 55,411 (FSU, GA, NY [fl], TENN, TEX).
 
Suwannee Co.: E of Lee off US 90 at Suwannee River State Park, 9 May 1988, Godfrey 82,666 (FSU).
 
Taylor Co.: SW of Cabbage Grove where Aucilla River is underground, 30 Jul 1957, Godfrey 55,788 (FSU [fr]).
 
Wakulla Co.: E of Wakulla along St. Marks River, 25 Mar 1955, Godfrey & Reed 53,050 (FSU, GA, NY).
 
Walton Co.: 7.1 mi E of Walton—Okaloosa Co. line by FL rte. 20, 18 Sep 1982, Godfrey 80,012 (FSU [fr]).
 
Washington Co.: along Choctawhatchee River nr. Caryville, 12 Jul 1969, Godfrey 68,766 (FSU [fr], GA).
 
Georgia
 
Baker Co.: Woodruff Plantation along Ichawaynochaway Creek N of Bridge on US 91, 19 Jul 1966, Chester & Chester 1252 (TENN [fr]), along Ichawaynochaway Creek behind Field Station, 17 Aug 1946, Thorne 1939 (GA, NY [fr], TENN), along Ichawaynochaway Creek nr. Field Station, 8 Apr 1947, Thorne & Muenscher 2840 (F [fl], US).
 
Baldwin Co.: nr. Old Fort Wilkinson along right bank of Oconee River, 14 Aug 1884, Smith s.n. (NY [fr], US).
 
Banks Co.: nr. Habersham Co. line along middle fork of Broad River, 31 May 1942, Duncan 5278 (GA).
 
Bartow Co.: 2.75 mi S of Allatoona Dam along W side of "elbow bend" of Allatoona Creek, 24 Apr 1948, Duncan 8160 (GA [fr], MO).
 
Bulloch Co.: 7 mi SW of Statesboro on rte. 25, 16 May 1953, Wilbur 3281 (FSU).
 
Burke Co.: 14.5 mi E of Waynesboro at bottom of High Head, 10 May 1990, Jones et al. 25,237 (GA).
 
Butts Co.: SE of Jackson off hwy. 42 in Indian Springs State Park, 25 May 1987, Howel & Jones 341 (GA).
 
Carroll Co.: Whooping Creek Rd. at Whooping Creek, 4 Apr 1977, Byrd 452 (GA [fl]).
 
Charleton Co.: Folkston, 15 May 1918, Harbison 14,302 (NCU [fr]).
 
Chatham Co., Savannah, 26 Mar 1867, Coultas s.n. (NY).
 
Clarke Co.: River Road on university campus, 16 Apr 1923, Miller E1348 (GA [fl]), Athens along Oconee River, 6 Oct 1979, Smith 747 (NY [fr], TEX), along Oconee River, 9 Apr 1978, Wolfe 33 (TENN [fl]).
 
Cobb Co.: N of Atlanta at Camp Bert Adams, 12 Aug 1941, Duncan 3910 (GA [fr]).
 
Dawson Co.: 2 mi S of Lumpkin Co. line on W side of Chestatee River, 1 May 1955, Duncan & Adams 18,538 (GA).
 
Decatur Co.: along US 27 3 mi S of Attapulgus, 19 Jul 1966, Chester & Chester 1040 (TENN [fl]), 1.6 mi NW of Amsterdam along Amsterdam Creek, 19 Aug 1965, Faircloth 2779 (GA [fr], MO, NCU), W side of Spring Creek at Brinson on US hwy. 84, 13 Mar 1976, Faircloth 7944 (NCU [fl]).
 
De Kalb Co.: along Peavine Creek in Druid Hills area of Atlanta, 9 Jul 1956, Burbanck 1538 (FSU, GA, LL, NCU, NY, TENN [fr]).
 
Dougherty Co.: along the Flint River at Albany, 24—28 May 1895, Small s.n. (NY [fr]), US).
 
Early Co.: along the Chattahoochee River at Saffold, 5 Apr 1947, Thorne & Muenscher 2753 (GA, MO [fl]), along Coheelee Creek nr. Hilton, 29 Oct 1947, Thorne 7412 (GA [fr]).
 
Effingham Co.: nr. Cedar Bluff along Savannah River, 31 Mar 1940, Eyles 6782 (GA [fl], TENN).
 
Elbert Co.: along Little Dove Creek, 3 Apr 1977, Coile et al. 435 (GA [fl]), beside Broad River above GA hwy. 172, 26 Jul 1980, Coile & Coile 2158 (GA [fr]).
 
Fannin Co.: 5 May 1926, Ashe s.n. (NCU [fl]), between Rich Knob and Crenshaw Gap along GA hwy. 2, 28 Apr 1979, Jones et al. 23,197 (F [fl], GA, TENN), Blue Ridge Mountains, 21 Jul 1909, Smith 2409 (F [fr]).
 
Floyd Co.: along Coosa River ca. 2 mi SW of Rome, 25 Mar 1945, Dorsey s.n. (GA [fl]), 3 mi SW of Rome along S side of Catoosa River, 26 Jun 1948, Duncan 8446 (GA [fr]), Mount Berry, 9 Apr 1939, Jones 165 (GA [fl]).
 
Forsyth Co.: Sawnee Mountain, 21 Apr 1956, Duncan 20,053 (GA [fl]).
 
Fulton Co.: S of Chattahoochee River and SE of Roswell, 4 Apr 1949, Duncan 9120 (GA [fl]), at Fulton—Coweta co. line along Chattahoochee River, 11 Apr 1937, Eyles 1418 (GA [fl]).
 
Gilmer Co.: 1 mi N of Potato Patch Mountain, 19 Sep 1946, Duncan 7228 (GA [fr]).
 
Glynn Co.: St. Simon’s Island, 17 Sep 1921, Harbison 15,983 (NCU).
 
Grady Co.: Effano Tobacco farm, 7 mi S of Whigham, 19 Mar 1964, Faircloth 750 (GA, NCU [fl]).
 
Gwinnett Co.: along Yellow River nr. McGuire’s Mill, 11 Jul 1893, Small s.n. (F, MO, US [fr]), 2 Aug 1895, Small s.n. (F [fr], MO, NY).
 
Habersham Co.: between Clayton and Tallulah Falls, 29 Aug 1932, Coker et al. s.n. (NCU [fr]), Tallulah Falls, Sep 1932, Harbison s.n. (NCU [fr]).
 
Hall Co.: 5 mi N of Gainesville in woods overlooking Little River, 11 May 1941, Duncan 3213 (GA [fr]).
 
Haralson Co.: Tallapoosa, Apr—May 1900, Way 48 (US).
 
Harris Co.: Pine Mountain, 15 Apr 1960, Henry 7212 (GA [fl]).
 
Hart Co.: W of Hartwell along Savannah River, 26 Apr 1942, Duncan 4843 (GA).
 
Jackson Co.: 2.5 mi SW of Braselton along Lee Creek, 6 Aug 1985, Boring & Lott 1728 (GA [fr]), 7.75 mi N of Jefferson on N branch of Oconee River along S bank, 4 Jul 1967, Underwood 68 (GA [fr]).
 
Jones Co.: nr. Buttlers Creek and Ocmulgee River junction, 21 Mar 1992, Payne & Payne s.n. (GA [fl]).
 
Long Co.: Glennville Township at Observation point 55, 22 Oct 1992, Nordman s.n. (GA).
 
Madison Co.: along Fork Creek NE of Carlton, 26 Apr 1942, Duncan 4786 (GA [fl]).
 
Monroe Co.: 1.5 mi off I—75 at exit 65 on High Falls Rd. in High Falls State Park, 12 Apr 1987, Howel 88 (GA [fr], MO).
 
Morgan Co.: 20 yd downstream from Lake Brantley Dam along Hard Labor Creek, 16 Apr 1980, Hill 1338 (GA, NCU, NY).
 
Murray Co.: Alaculsy Valley along GA hwy. 2, 17 Apr 1980, DiGioia 264 (GA [fl], NCU), between Lake Conasauge and Potato Patch Mountain along Forest Service Rd. 68, 29 Aug 1979, Jones et al. 23,546 (GA, NCU, NY [fr]).
 
Oconee Co.: floodplain of Oconee River at Barnett Shoals Bridge, 12 Aug 1967, Jones 15,045 (LSU [fr]).
 
Oglethorpe Co.: along Buffalo Creek E of Lexington, 18 Apr 1942, Duncan 4757 (GA [fl]).
 
Pickens Co.: 2 mi SW of Marble Hill nr. junction of Cox’s Creek and East Creek, 8 Sep 1988, Jones et al. 25,066 (GA [fr]).
 
Rabun Co.: nr. Wylie by stream, 5 Sep 1933, Totten s.n. (NCU [fr]).
 
Randolph Co.: Anderson, 5 Apr 1914, Harbison 11,457 (NCU).
 
Richmond Co.: nr. Augusta, 31 Mar 1913, Harbison s.n. (GA [fl]), 4 Apr 1914, Harbison 11,450 (NCU [fl]), 4 Apr 1914, Harbison (1)1451 (NCU [fl]), 4 Apr 1914, Harbison (1)1452 (NCU [fl]), 4 Apr 1914, Harbison 11,455 (TENN [fl]).
 
Screven Co.: E of Newington and S of Blue Spring along small stream, 7 Aug 1942, Duncan 5591 (GA [fr]).
 
Seminole Co.: Sealy’s Plantation, 26 Mar 1947, Thorne & Muenscher 2427 (GA, NY).
 
Stephens Co.: 2 mi up Panther Creek from junction with Tugaloo River, 16 Apr 1978, Graves s.n. (GA [fl]), along Panther Creek 1.3 mi SW of junction with Tugaloo River, 12 May 1976, Solomon 1764 (MO [fr]).
 
Sumter Co.: 15.2 mi E of Sumter Co. Courthouse along Flint River, 19 Aug 1965, Parker 109 (GA).
 
Tattnall Co.: E of Ohoopee River along US 280, 21 Aug 1940, Duncan 2704 (GA).
 
Taylor Co.: along Flint River NE of Butler, 28 Mar 1953, Duncan & Hardin 14,857 (GA [fl]).
 
Thomas Co.: 3.3 mi E of Ochlocknee along E bank of Little Ochlocknee River, 4 Apr 1964, Faircloth 787 (NCU [fl]).
 
Towns Co.: Tray Mountain, Mossy Cove, 1 May 1986, Chafin 522 (GA [fl]).
 
Union Co.: N slopes of Sassafras Mountain, 12 Jul 1942, Duncan 5481 (GA [fr]).
 
Upson Co.: 6 mi S of Thomaston, 18 Apr 1959, Godfrey 58,423 (FSU [fl], NY).
 
Walton Co.: nr. Morgan Co. line along bottomland of Jacks Creek, 3 Nov 1946, Duncan 7379 (GA [fr]).
 
Wayne Co.: Island Maritime Forest, S side Altamaha River, 9 Jul 1993, Pyne 93—170 (TENN).
 
White Co.: ravine along N side of Yonah Mountain, 7 May 1966, Duncan 22,673 (GA [fr]).
 
Wilkes Co.: along Broad River at Anthony Shoals, 26 Aug 1939, Duncan 1626 (GA [fr]).
 
Worth Co.: 2 mi WSW of Warwick below hydroelectric dam on Flint River, 17 Mar 1967, Faircloth 4111 (GA, NCU).
 
Illinois
 
Massac Co.: along Ohio River hillsides nr. Metropolis, 9 May 1923, Palmer 22,573 (F, MO [fl]), Metropolis along Ohio River, 15 Sep 1923, Palmer 23,742 (GA, MO [fr], US).
 
Indiana
 
Perry Co.: Canelton, 1848, Short s.n. (MO).
 
Vanderburgh Co.: bank of Ohio River nr. Evansville, 1848, Short s.n. (MO [fr]).
 
Kentucky
 
Lawrence Co.: Louisa, 1837, Short s.n. (TEX).
 
Marshall Co.: 2 mi S of Tennessee River between Calvert City and Paducah, 22 Sep 1945, Duncan 6186 (GA [fr], MO).
 
McCracken Co.: along Tennessee River, 2 May 1937, Fleetwood s.n. (NY [fl]).
 
Louisiana
 
Rapides Co.: E of Flatwoods on hwy. 8, 5 Apr 1969, Brown 20,318 (LSU).
 
Mississippi
 
Jones Co.: Laurel, 26 Mar 1917, Harbison 13,029 (TENN).
 
Lauderdale Co.: S of Chunky and I—20 along Allen Creek, 12 Jul 1978, Rogers 45,309 (TENN).
 
Perry Co.: 2 mi S of Runnelstown along Tallahala Creek, 22 Mar 1967, Jones 11,221 (FSU, GA).
 
Wayne Co.: 13 mi NE of Waynesboro along S18 junction with Shiloh Creek S of Matherville, 19 Mar 1977, McDaniel 21,172 (TEX [fl], NY).
 
North Carolina
 
Anson Co.: Blewett Falls on Pee Dee River, 30 Jun 1956, Radford 13,325 (NCU).
 
Ashe Co.: without date and locality, Ashe s.n. (NCU).
 
Avery Co.: Lower Grandfather Mountain, 14 Aug 1890, Sudworth s.n. (US).
 
Bladen Co.: "Paradise" Arboretum at Flora McDonald College (brought in from between Elizabethtown and Tar Heel), 18 Apr 1937, Totten s.n. (NCU).
 
Buncombe Co.: nr. Asheville, 18 Apr 1929, Palmer 35,419 (MO, US [fl]), Biltmore, 24 Apr 1896, Biltmore Herbarium No. 520 (F [fl], NY, US), 27 Apr 1897, Biltmore Herbarium No. 520c (A, F, NCU, NY, US [fl]).
 
Burke Co.: nr. Hawkbill Mountain lookout by old NC 105, 9 Sep 1958, Bell 15,154 (NCU [fr]), 0.5 mi SE of Table Rock, 1 May 1953, Radford et al. 6883 (US [fl]), Table Rock Mountain, 6 Sep 1972, Thomas 242 (NY [fr]).
 
Caldwell Co.: 25 Apr 1955, Hyler et al. s.n. (NCU [fl]), 3.5 mi E of Oak Hill nr. NC 90, 4 Aug 1956, Radford 15,091 (NCU [fr]).
 
Catawba Co.: nr. Hickory, 18 Sep 1938, Ivey s.n. (NCU [fr]), 3.5 mi upriver from NC hwy. 150 bridge along floodplain on SW side of Catawba River, 17 Aug 1959, McNeely 381 (NCU [fr]).
 
Cherokee Co.: Andrews by Valley River, 8 Sep 1932, Harbison & Totten s.n. (NCU [fr]), along a tributary of the Valley River E of Andrews along rte. 19, 24 Apr 1986, Hill 16,640 (NY [fl]).
 
Clay Co.: 2.25 mi N of Hayesville along Hiwassee River, 5 Jun 1952, Radford & Wood 6147 (NCU, TEX).
 
Cleveland Co.: 4 mi S of Boiling Springs nr. rte. 150 along Broad River, 5 Aug 1947, Fox 674 (GA [fr]).
 
Cumberland Co.: 6 mi below Fayetteville along Cape Fear River, 26 Sep 1936, Coker & Rankin s.n. (NCU, NY [fr]), 8 Jun 1936, Rankin s.n. (NCU [fr]).
 
Davidson Co.: rocky hillside above "Little Cliff" on Yadkin River, 16 Aug 1931, Totten s.n. (NCU [fr]), 17 Aug 1931, Totten s.n. (NCU [fr]).
 
Davie Co.: Yadkin River nr. US 64, 14 Apr 1956, Radford 9808 (NCU, NY [fl]).
 
Durham Co.: Aunt Sallie's Ridge N of Piedmont Springs, 1—3 Jul 1891, Small and Heller 289 (F, MO, NY, US).
 
Forsyth Co.: nr. Winston-Salem, 20 Apr 1957, Schallert 654 (FSU).
 
Gaston Co.: 1 mi S of Mountain View nr. Crowder’s Mountain, 21 Jun 1956, Ahles & Leisner 15,067 (NCU).
 
Graham Co.: nr. Robbinsville, 5 Oct 1942, Stearns s.n. (US [fr]).
 
Haywood Co.: Canton, 3 May 1924, Knowlton s.n. (NCU [fl]), lower slopes of Mount Pisgah, 6 May 1917, Maxon 6297 (US [fl]), W fork of Pigeon River, 8.5 mi N of Beech Gap, 2 May 1953, Radford & Wood 6900 (NCU [fl]), along a stream nr. Waynesville, 10 Sep 1910, Standley 5725 (US [fr]).
 
Henderson Co.: rd. to Flat Rock by meadow, 22 Aug 1910, Coker s.n. (NCU [fr]), Bat Cave, 16 Apr 1973, Moldenke & Moldenke 27,071 (LL [fl]), 5 mi NE of Hendersonville, 0.5 mi W of US 64, 6 May 1956, Pittillo 59 (NCU [fl]).
 
Iredell Co.; 0.4 mi S of NC 150 along Catawba River, 1 Jul 1958, Ahles & Duke 45,110 (NCU).
 
Jackson Co.; NC 107, 7.3 mi S of junction with NC 281, 2 May 1965, Bernstein s.n. (NCU [fl]), nr. Wet Camp Gap, without date, Bond 21 (NY [fl]), Balsam, 16 Nov 1911, Braun s.n. (US [fr]), roadside at White Side Cove, 16 Sep 1932, Harbison & Totten s.n. (NCU [fr]), Pisgah National Forest 7 mi E of Blue Ridge Parkway junction with US 441, 22 May 1983, Hess 5837 (GA, NCU [fl]), rte. 107 ca. 5 mi N of Cashiers, 2 May 1991, Hill 22,190 (MO [fl], NCU, NY), along rd. to Upper Whitewater Falls, 13 Sep 1976, Rodgers & Mullens 76,532 (MO [fr]).
 
Macon Co.: 22 Apr 1911, Harbison 506 (A, F, MO, US [fl]), 1.5 mi below Cullasaja Falls in pasture on Peeks Creek Rd., 25 Jul 1938, Hechenbleikner & Stewart s.n. (NCU [fr]), on hill of old home site back of Episcopal Church, 23 Aug 1944, Stewart s.n. (NCU [fr]), along Buck Lake Rd. at Rattlesnake Rd., 7 May 1970, Summerlin 147 (TENN [fl]), bank of Cullosajo River ca. 10 mi NW of Highlands, 12 Aug 1948, Wilbur 1221 (TEX [fr]).
 
Madison Co.: French Broad River E of Paint Rock, 22 May 1966, Chester 1146 (TENN).
 
McDowell Co.: 0.1 mi E of Buncombe Co. line ca. 5.5 mi W of Old Fort, 20 Apr 1956, Ahles & Bell 11,137 (NCU [fl]), 12 Aug 1938, Rogers 8459 (MO [fr], NY, TEX, US).
 
Mecklenburg Co.: bluff of Catawba River, 6 Apr 1953, Daggy 2142 (NCU).
 
Montgomery Co.: along Yadkin River below Narrows Dam, 9 May 1970, Wells 2839 (NCU).
 
Polk Co.: base of Ballingers Hill, 20 Jun 1921, Peattie 606 (F [fr]), nr. Columbus, 14 Apr 1897, Townsend s.n. (US [fl]).
 
Rowan Co.: SW of US 601 bridge across South Yadkin River, 7 Apr 1957, Horton 654 (NCU).
 
Rutherford Co.: along Cane Creek Rd., 2 mi ENE of US 64, 21 Apr 1956, Bell 2096 (NCU [fl]).
 
Swain Co.: along rte. 19, 5.4 mi SW of rte. 28—W junction along the Nantahala River, 24 Apr 1986, Hill 16,654 (MO, NCU [fl], NY), Hickory Nut Gap, 3 May 1902, Hubbard s.n. (LL [fl]), High Rock, 11 Sep 1935, Jennison 783 (TENN [fr]), nr. New Found Gap, 4 May 1936, Jennison 2279 (TENN [fl]), 1.5 mi SE of Alarka, 24 Aug 1956, Radford 16,941 (TENN [fr]), 0.5 mi below Board Camp Trail along Forney Creek Rd., 13 Aug 1939, Stewart & Howell s.n. (NCU [fr]), Wesser along Appalachian Trail, Jul 1976, Williams s.n. (NY [fr]).
 
Transylvania Co.: on hwy. 28 nr. Lake Toxaway, 8 Aug 1932, Coker s.n. (NCU [fr]), 0.5 mi N of rte. 276 on Forest Rd. 1206, Pisgah National Forest, 29 Apr 1989, Hill 20,506 (NY [fl]), "Pink Beds", 10 May 1909, House 4133 (US [fl]), 5.9 mi S of Blue Ridge Parkway on NC 215, 9 May 1980, Wood et al. 4558 (MO [fl], NY).
 
Wake Co.: Raleigh, 20 Sep 1924, Ashe s.n. (NCU).
 
Watauga Co.: Blowing Rock, 8 Aug 1890, Sudworth 77 (US).
 
Wilkes Co.: 1 mi NW of Ball’s Mill site, 3 May 1941, Radford & Stewart 1916 (GA, NCU [fl]).
 
Yancey Co.: at Yancy—McDowell co. line, 16 May 1941, McQuilkin 8600 (TEX, US [fl]).
 
Ohio
 
Pike Co.: Chimney Rocks E of Omega Hillside and Bottomland of Scioto River, 16 Aug 1965, Johnson 147 (NCU).
 
Oklahoma
 
Le Flore Co.: 15 mi N of Smithville, 24 Apr 1952, Blinn 34 (TEX [fl]), top of Kiamichi Mountain, Jun 1927, Ortenburger s.n. (US [fr]), N side of Rich Mountain, 12 Apr 1922, Palmer 20,763 (NY [fl]), Le Flore—Pushmataha co. line, SW of Honobia along Little River, 3 Jul 1948, Robbins 3120 (NY [fr]), 25 Apr 1931, Stemen & Meyers s.n. (US [fl]).
 
McCurtain Co.: N of Hochatown at Mountain River, 26 Aug 1960, Sharp s.n. (TENN [fr]), 1.75 mi SW of Smithville along Eagle Fork Creek, 21 Jun 1977, Taylor & Taylor 24,668 (LSU [fr]).
 
Pushmataha Co.: 1.5 mi NW of Honobia along Honobia Creek, 20 Jun 1977, Taylor 24,634 (LSU).
 
South Carolina:
 
Abbeville Co.: 0.2 mi S of SC 72 bridge over Savannah River along Savannah River at Russell Dam and Lake area, 11 Apr 1978, Fairey 704 (GA, NCU, NY).
 
Aiken Co.: nr. Hazel’s Lake, Beech Island, 25 Mar 1946, Radford 1383 (NCU [fl]).
 
Anderson Co.: Andersonville, 1885, Gibbes s.n. (NY [fl]).
 
Beaufort Co.: Bluffton, May 1882, Mellichamp s.n. (NY).
 
Berkeley Co.: Woodlawn Plantation (Mr. S. S. DeVeawn's) St. John's, without date, Porcher s.n. (NY [fl]).
 
Calhoun Co.: SE of Lone Star on SC 267 at Halfway Swamp Creek, 3 Jul 1957, Ahles & Haesloop 30,259 (NCU).
 
Charleston Co.: La Roche Farm above Salt Marsh on John’s Island, 25 Mar 1944, Hunt & Martin 2446 (NCU).
 
Cherokee Co.: along Bowens River on co. Rd. 83 1 mi N of junction with SC 18, Ahles & Haesloop 30,964 (NCU [fr]).
 
Chester Co.: 0.5 mi SE of Lockhart nr. the Broad River, 7 Apr 1956, Freeman 5644 (NCU).
 
Dorchester Co.: banks of Edisto River in Givhans Ferry State Park, 14 Jun 1988, Hill 19,589 (NY).
 
Edgefield Co.: Sumter National Forest, 30 May 1993, Hill & Dorsey 25,075 (TEX).
 
Georgetown Co.: nr. Brookgreen Garden, 14 Sep 1940, Totten et al. s.n. (NCU [fr]).
 
Greenville Co.: Gap Creek Rd., River Falls, 7 Apr 1967, Mullens & Rodgers 67,036 (FSU, GA, LL, NY [fl], TENN, TEX).
 
Jasper Co.: 5.5 mi NW of Tillman nr. Savannah River, 14 Apr 1968, Leonard 1267 (NCU [fl, fr]).
 
Kershaw Co.: 8 mi NW of Camden along Sawneys Creek, 7 Apr 1957, Radford 20,841 (NCU).
 
Lancaster Co.: 1.1 mi SE of junction SC rte. 97 and rte. 200 on SC rte. 97 (SW of Lancaster), 15 Apr 1967, Ahles & Haesloop 23,076 (NCU).
 
Laurens Co.: 12 mi SW of Laurens along Reedy River at Boyd Mill Pond, 12 Apr 1957, Bell 5570 (NCU).
 
Lexington Co.: 1 mi S of Cayce along Congaree River, 5 Apr 1957, Radford 20,509 (NCU).
 
Oconee Co.: 24 Sep 1924, Ashe s.n. (NCU [fr]), 4 mi E of Mountain Rest along small creek, 11 Jul 1939, Harper 1340 (NCU [fr]), along Seneca River, 18 Apr 1906, House 1866 (MO, NY, US [fl]), riverbank of Devil’s Fork, 300 yd upstream from junction with Whitewater River, 26 Aug 1965, Powell & Patton 65—102 (NCU [fr]).
 
Pickens Co.: Old Hagwood Mill N of Pickens, 26 Sep 1991, Hill 22,644 (NY [fr]), nr. SC hwy. 183 W of Pickens along Kegwee River, 22 Aug 1956, Radford 16,419 (NCU [fr]), W of Pickens along Twelve Mile Creek nr. SC hwy. 183, 22 Aug 1956, Radford 16,675 (NCU [fr]).
 
Richland Co.: Columbia, May 1890, Taylor s.n. (F).
 
Saluda Co.: Saluda, 24 Oct 1916, Harbison s.n. (NCU).
 
Spartanburg Co.: N of Cross Anchor at Tyger River, 13 Apr 1957, Bell 5634 (NCU).
 
Williamsburg Co.: at the Santee River Bridge ca. 2.7 mi N of Jamestown, 6 Apr 1975, Reveal & Seldin 3832 (F, FSU [fl], GA, GH, MO, NY, TEX, US).
 
Tennessee
 
Anderson Co.: E of Lake City on Coal Creek, 17 Apr 1966, Chester & Chester 1044 (TENN).
 
Blount Co.: Cades Cove Mountain, 13 Apr 1929, Anderson 1156 (NY [fl]), along old hwy. to Maryville nr. Walland, 17 Apr 1956, Brown s.n. (FSU [fl]), Cades Cove Mountain, 11 May 1933, Cain 78 (TENN [fl]), 11 May 1933, Duncan 429 (GA [fl]), above Rockford on steep slopes of Little River, 29 Sep 1963, Gaylon 32,249 (TENN [fr]), nr. Walland, 30 Apr 1934, Hudson s.n. (TENN [fl]), Cades Cove Mountain, 13 Apr 1929, Jennison & Anderson s.n. (MO [fl], US), along Little River nr. Kinzel Springs, 22 Apr 1934, Jennison 308 (TENN [fl]), Townsend, 1 May 1932, Kinsey s.n. (TENN [fl]), below Tallassee below Chilhowee Dam site, 29 Apr 1956, Sharp & Gaylon 20,475 (NY [fl], TENN), Miller’s Cove along Rich Creek, 29 Apr 1961, Sharp et al. 28,108 (GA [fl], TENN), Rich Mountain, 22 Apr 1934, Underwood & Sharp 510 (GA [fl], TENN).
 
Cocke Co.: Hwy. 32, 3 mi E of Crosby, 7 Apr 1977, Barton 26 (LSU [fl]), along hwy. 107 1.7 mi W of junction with 25—70, 12 Sep 1965, Chester 952 (TENN [fr]), along French Broad River between Paint Rock and Del Rio, within 3 mi of Wolf Creek Station, 24 Aug 1897, Kearney 827 (MO [fr], NCU, NY, US [fl]), nr. Del Rio along Newport Rd., 18 Apr 1953, Sharp et al. 17,252 (TENN [fl]), Pigeon River bluff below Watertown, 29 Jul 1970, Sharp & Bowers 44,484 (TENN [fr]).
 
Decatur Co.: S of Mount Carmel Church along Tennessee River, 14 Oct 1949, RES (Shanks) 14,504 (TENN).
 
Greene Co.: Near Point Creek nr. French Broad River, 21 May 1966, Sharp et al. 41,037 (TENN).
 
Hamilton Co.: at border of Hamilton Co. & Walker Co., Georgia, Lookout Mountain, 18 Apr 1906, Churchill s.n. (TENN [fl]), Elise Chapin Sanctuary, 26 Jul 1952, RES (Shanks) 16,871 (TENN).
 
Humphreys Co.: branch of Tennessee River off I—40, 7.5 mi W of Buffalo, 24 Apr 1972, Kral 45761 (MO).
 
Knox Co.: 2 mi W of Knoxville on S side of Tennessee River, 1 Aug 1965, Chester 933 (TENN [fr]), Tennessee River bank nr. Knoxville, 23 Apr 1966, Chester 1046 (TENN [fl]), Peter Blow farm beyond Maxey’s Bluff, 25 Apr 1956, Gaylon 20,438 (TENN [fl], FSU), nr. Knoxville, Apr 1893, Kearney s.n.(US [fl]), Jul 1892, Ruth s.n. (MO [fr]), Tennessee River bank nr. Knoxville, Jul 1899, Ruth 3020 (NCU [fl], NY), nr. Knoxville, 20 Apr 1889, Scribner s.n. (NY [fl]), Tennessee River bank Island Home Knoxville, 25 Apr 1931, Wilson s.n. (TENN [fl]).
 
Loudon Co.: overlooking Tennessee River by Tellico Dam, 17 Jun 1990, McNeilus 90—386 (TENN).
 
Marion Co.: McGlothlin Tract #3 US hwy. 41/64/72 ca. 3 mi NE of Haletown, 12 Apr 1994, Pyne et al. 94—029 (TENN [fl]), Sequatichie Valley N of Jasper, 22 Apr 1951, Shanks et al. 15,458 (FSU, GA, LL, MO, NCU [fl], NY, TENN, US).
 
Monroe Co.: 1 mi S of Tellico Mountain Camp along hwy. 68, 6 Oct 1965, Chester et al. 977 (TENN [fr]), Citico Creek along state line, 30 Apr 1977, Malter 53,094 (TENN [fl]), nr. Beech Gap in Cherokee National Forest, 30 May 1940, Sharp 699 (TENN [fl]).
 
Polk Co.: 1 mi upstream from Appalachian Powerhouse along the N shore of the Hiawassee River, 18 Apr 1980, Manek 85 (TENN [fl]), Copperhill, 20 Jul 1969, Rogers 44,060 (TENN [fr]), above Hiawassee River N of bridge across hwy. 68, 23 Apr 1966, Rogers & Rogers 44,926 (NCU [fl]).
 
Roane Co.: Watts Bar Lake at river mi 563 W of Chapman Branch, 14 Jun 1984, Wofford 84—38 (TENN).
 
Sevier Co.: Elkmont, 28 Sep 1924, Ashe s.n. (NCU [fr]), on Pigeon Forge-Walland Rd. ca. 2 mi from Blount Co. line, 19 Jul 1965, Chester 926 (TENN [fr]), W prong of Pigeon River, 27 Apr 1966, Chester 1085 (TENN [fl]), Blanket Mountain Trail in Elkmont, 5 May 1957, Comte 1120 (MO [fl]), 13 Apr 1934, Duncan 652 (GA [fl]), nr. Gatlinburg along Little Pigeon River, 27 Apr 1931, Haynie 2233 (F [fl]), on Cherokee Orchard Rd. above Gatlinburg, without date, Jennison s.n. (TENN [fl]), nr. Gatlinburg nr. Great Smoky Mountains National Park boundary, 21 Apr 1936, Jennison 2152 (TENN [fl]), 6 May 1947, Miller 2123 (US [fl]), nr. Gatlinburg, 12 Oct 1944, Miller 2222 (US [fr]), edge of Carrs Motel in Gatlinburg, 26 Apr 1969, Whatley 158 (NCU [fl]).
 
Stewart Co.: N of Ginger Creek embayment of Kentucky Lake (Tennessee River), 4 Jun 1966, Chester & Lyle 1177 (TENN).
 
Unicoi Co.: Unaka Springs nr. Erwin, 5 May 1934, Cook 24 (TENN [fl]).
 
Washington Co.: 0.5 mi N of Embreville Bridge along Nolichucky River, 20 Jun 1965, Chester 877 (TENN).
 
Virginia
 
Carroll Co.: along New River on Farmer Mountain, 12 Jul 1892, Small s.n. (F [fr], NY).
 
Giles Co.: nr. Norris Run, 28 Apr 1961, Kral 12,210 (NCU [fl]).
 
Grayson Co.: 2.5 mi W of Galax Corp. limits along hwy. 58 along New River, 9 May 1976, Wieboldt & Stevens 2419 (NCU).
 
Montgomery Co.: 2 mi SSW of Goodwin Ferry Station S of Giles Co. line, 16 Aug 1956, Thorne 18,147 (FSU).
 
Pulawski Co.: along the New River, 1878, Shriver s.n. (US [fl]).
 
Wythe Co.: Wytheville, 29 Apr 1874, Shriver s.n. (NY, US).
 
West Virginia.
 
Fayette Co.: nr. Glenn Ferris, 16 Apr 1953, Bartley 1945 (US [fl]), banks of Kanawha River, 18 Jul 1890, Millspaugh 601 (NY [fr]), Quinnemont, 21 Aug 1899, Pollard & Maxon s.n. (NY [fr]).
 
Kanawha Co.: along hwy. 61 between Montgomery and Handley, 12 Apr 1973, Adkins 2826 (GA).
 
Raleigh Co.: NW side of New River directly above Sandstone Falls, 12 Aug 1984, Cusick & Spooner 23,802 (NCU [fr]).
 
Summers Co.: Haynes Ferry, 16 Jul 1930, Berkley 1627 (MO [fr]), 15 Apr 1967, McNeill s.n. (NCU [fl]).