Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Gasterosteus: [ 53 ] records

aculeatus, Gasterosteus Linnaeus [C.] 1758:295 [Systema Naturae, Ed. X v. 1; ref. 2787] Europe. Syntypes: Zool. Soc. Lond. 29 (left half-skin), 30-31 (2, right half-skins). •Valid as Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363], Kawanabe & Mizuno 1989:433 [ref. 25598], Pietsch et al. 2001:144 [ref. 25688], Moyle 2002:340 [ref. 26449], Scharpf 2007:34 [ref. 30398]). •Valid as Gasterosteus aculeatus Linnaeus 1758 -- (Herrera 1896:38 [ref. 39336], Berg 1949:972 [ref. 20662], Monod 1973:280 [ref. 7193], Lee et al. 1980:563 [ref. 22416], Eschmeyer & Herald 1983:128 [ref. 9277], Ida in Masuda et al. 1984:83 [ref. 6441], Smith 1986:276 [ref. 22186], Banister in Whitehead et al. 1986:640 [ref. 13676], Arnoult 1986:280 [ref. 6213], Robins & Ray 1986:122 [ref. 23100], Lelek 1987:267 [ref. 22962], McAllister 1990:217 [ref. 14674], Page & Burr 1991:243 [ref. 18983], Haglund et al. 1992 [ref. 19890], Espinosa Pérez et al. 1993:62 [ref. 22290], Swift et al. 1993:123 [ref. 22379], Jenkins & Burkhead 1994:623 [ref. 21581], Poll & Gosse 1995:109 [ref. 24781], Coad 1995:26 [ref. 23608], Zhu 1995:165 [ref. 25213], Mrakovcic et al. 1995:185 [ref. 23079], Chereshnev 1996:605 [ref. 24659], Coad 1996:85 [ref. 25304], Bianco et al. 1996:56 [ref. 26724], Cheng & Zhou 1997:233 [ref. 26385], Kottelat 1997:164 [ref. 22952], Murdy et al. 1997:135 [ref. 23144], Kim 1997:386 [ref. 24032], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:747 [ref. 24702], Bianco 1997:620 [ref. 26725], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Caldecutt & Adams 1998:827 [ref. 23613], Ahnelt et al. 1998:395 [ref. 23708], Scott & Crossman 1998:665 [ref. 24570], Sokolovskaya et al. 1998:9 [ref. 24670], Castro-Aguirre et al. 1999:226 [ref. 24550], Fuller et al. 1999:334 [ref. 25838], Lyons et al. 2000:51 [ref. 26594], Nakabo 2000:513 [ref. 25086], Sheiko & Fedorov 2000:25 [ref. 25650], Bogutskaya et al. 2001:46 [ref. 26178], Caldecutt et al. 2001:936 [ref. 25815], Chereshnev et al. 2001:112 [ref. 26736], Zemnukhov et al. 2001:420 [ref. 25412], Bilecenoğlu et al. 2002:64 [ref. 26753], Kim & Park 2002:308 [ref. 25913], Krueger in Collette & Klein-MacPhee 2002:314 [ref. 26158], Mecklenburg et al. 2002:333 [ref. 25968], Nakabo 2002:513 [ref. 26001], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Roe et al. 2002:903 [ref. 26504], Choi et al. 2003:231, 552 [ref. 26218], Pichugin et al. 2003:169 [ref. 26890], Hagiwara 2003:37 [ref. 27188], Fedorov et al. 2003:57 [ref. 27579], Jang et al. 2003:120 [ref. 27681], Wydoski & Whitney 2003:167 [ref. 27816], Hanel 2003:61 [ref. 27894], Alekseyev et al. 2004:54 [ref. 27625], Bailey et al. 2004:168 [ref. 27882], Nelson et al. 2004:111 [ref. 27807], Bogutskaya & Naseka 2004:180 [ref. 28183], Love et al. 2005:71 [ref. 37547], Vassilev & Pehlivanov 2005:175 [ref. 28369], Miller 2006:320 [ref. 28615], Hanel & Lusk 2005:334 [ref. 28804], Fricke 2007:28 [ref. 30577], Fricke et al. 2007:73 [ref. 29533], Kottelat & Freyhof 2007:492 [ref. 29996], Vasil'eva 2007:70 [ref. 30517], McPhail 2007:432 [ref. 31199], Ninua & Japoshvili 2008:169 [ref. 31251], Kuiter 2009:322 [ref. 30404], Esmaeili et al. 2010:375 [ref. 33391], Lipej & Dulčić 2010:36 [ref. 36649], Møller et al. 2010:50 [ref. 31169], Mecklenburg et al. 2011:121 [ref. 31212], Page & Burr 2011:464 [ref. 31215], Page et al. 2013:113 [ref. 32708], Bianco 2013:9 [ref. 32897], Parin et al. 2014:203 [ref. 33547], Jouladeh-Roudbar et al. 2015:895 [ref. 34062], Pietsch & Orr 2015:29 [ref. 34694], Çiçek et al. 2015:151 [ref. 35151], Denys et al. 2015:41 [ref. 35420], Afonso et al. 2016:39, 155 [ref. 36199], Kells et al. 2016:122 [ref. 35888], Zhang et al. 2016:192 [ref. 34477], Dyldin & Orlov 2017:56 [ref. 35138], Esmaeili et al. 2017:95 [ref. 35254], Saygun et al. 2017:187 [ref. 38891], Esmaeili et al. 2018:77 [ref. 36089], Artüz & Fricke 2019:555 [ref. 36612], Burton & Lea 2019:56 [ref. 37205], Ćaleta et al. 2019:190 [ref. 37026], Carneiro et al. 2019:297 [ref. 37250], Lorenzoni et al. 2019:252 [ref. 37108], Özuluğ & Saç 2019:27 [ref. 36599], Ahnelt et al. 2020:105 [ref. 39966], Çiçek et al. 2020:259 [ref. 37644], Dyldin et al. 2020:85 [ref. 37569], Jouladeh-Roudbar et al. 2020:284 [ref. 37983], Kim et al. 2020:85 [ref. 39118], Kovačić et al. 2020:37 [ref. 37519], Kuljanishvili et al. 2020:[5] [ref. 37535], Abliazov et al. 2021:11 [ref. 38576], Bañón & Maño 2021:65 [ref. 38883], Dyldin et al. 2021:58 [ref. 38804], Love et al. 2021:102 [ref. 39279], Denys et al. 2022:396 [ref. 40085], Dyldin & Orlov 2022:35 [ref. 38972], Niță et al. 2022:48 [ref. 40471], Simian et al. 2022:375 [ref. 40776], Çiçek et al. 2023:450 [ref. 40625], Mousavi-Sabet et al. 2023:35 [ref. 40667], Saad et al. 2023:47 [ref. 40501], Shumka et al. 2023:38 [ref. 40747], Çiçek et al. 2024:375 [ref. 40789]). Current status: Valid as Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Distribution: North Pacific; Atlantic; Baltic Sea; North Sea; Mediterranean Sea; Sea of Marmara; Black Sea and adjacent watersheds; Sea of Japan; widespread in northern Europe, northern Asia and North America. Introduced in Caspian Sea region. Habitat: freshwater, brackish, marine.

algeriensis, Gasterosteus Sauvage [H.-E.] 1874:17 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] Algeria. Syntypes: MNHN 0000-5123 (4). Type catalog: Bertin & Estève 1950:38 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Arnoult 1986:280 [ref. 6213], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

argentatissimus, Gasterosteus Blanchard [C. E.] 1866:232, Figs. 35-36 [Les poissons des eaux douces de la France; ref. 310] Brooks near Avignon, France. Syntypes: BMNH 1883.7.4.5-10 [ex MNHN] (6?); MNHN 0000-6675 (6), 6678 (8 + 10); USNM 21140 [ex MNHN] (5). Type catalog: Bertin & Estève 1950:38 [ref. 19576]. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

argyropomus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:498 [Histoire naturelle des poissons v. 4; ref. 998] Florence, Italy. Syntypes: MNHN 0000-0677 (3). Type catalog: Bertin & Estève 1950:35 [ref. 19576]. Appeared first as name only in Cuvier 1829:170 [ref. 995]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

atkinsii, Gasterosteus Bean [T. H.] 1879:67 [Proceedings of the United States National Museum v. 2 (no. 71); ref. 16037] Schoodic Lakes, Maine, U.S.A. Syntypes: USNM 22492 (6, now 5). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:283 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

bailloni, Gasterosteus Blanchard [C. E.] 1866:231, Fig. 34 [Les poissons des eaux douces de la France; ref. 310] Abbeville, France. Syntypes: MNHN 0000-7151 (7), 0000-7153 (12). Type catalog: Bertin & Estève 1950:37 [ref. 19576]. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

biaculeatus, Gasterosteus Shaw [G.] 1803:608 [General zoology or systematic natural history v. 4; ref. 4014] New York, U.S.A. •Perhaps an unneeded new name for Gasterosteus bispinosus Walbaum 1792. Current status: Synonym of Gasterosteus. Gasterosteidae. Habitat: freshwater, brackish, marine.

biaculeatus, Gasterosteus Mitchill [S. L.] 1815:430, Pl. 1 (fig. 10) [Transactions of the Literary and Philosophical Society of New-York v. 1 (art. 5) (for 1814); ref. 13292] New York, U.S.A. No types known. Apparently not an original description but an identification of Gasterosteus biaculeatus Shaw. •In the synonymy of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Smith 1986:277 [ref. 22186], Denys et al. 2022:396 [ref. 40085]). 1°Homonym. Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

biarmatus, Gasterosteus Krynicki [J.] in Nordmann 1840:381 [Voyage dans la Russie méridionale et la Crimée; ref. 3209] Lagoons near Tarkanckut, Crimea. No types known. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:973 [ref. 20662], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Parin et al. 2014:203 [ref. 33547], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

bispinosus, Gasterosteus Walbaum [J. J.] 1792:450 [Petri Artedi sueci genera piscium Part 3; ref. 4572] New York, U.S.A. Apparently an unneeded new name for Gasterosterus bispinosus Walbaum 1792. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:280 [ref. 7193], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

brachycentrus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:499, Pl. 98 (fig. 2) [Histoire naturelle des poissons v. 4; ref. 998] Florence, Italy. Syntypes: MNHN 0000-0694 (3), 0000-7097 (2); SMF 1181 [ex MNHN] (1). Type catalog: Bertin & Estève 1950:35 [ref. 19576]. Appeared first as name only in Cuvier 1829:170 [ref. 995]. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

cataphractus, Gasteracanthus Pallas [P. S.] 1814:229 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Kamchatka, Russia. •Valid as Gasterosteus aculeatus Linnaeus 1758, variety cataphractus Pallas 1814 -- (Herrera 1896:38 [ref. 39336] as cataphractos). •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:972 [ref. 20662], Monod 1973:281 [ref. 7193], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Mecklenburg et al. 2002:333 [ref. 25968], Parin et al. 2014:203 [ref. 33547], Dyldin & Orlov 2017:56 [ref. 35138], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

crenobionta, Gasterosteus aculeatus forma Băcescu [M. C.] & Mayer [R.] 1956:24, Figs. 2b, 3c, e, 4b, d [Buletinul Institutului de Cercetari Piscicole. v. 15 (no. 2); ref. 21859] Lake Techirghiol, Romania. Syntypes: MGAB 49912a-b (2). Type catalog: Mihai-Bardan 1984:460 [ref. 21542] with a holotype. Apparently acceptable as a forma = subspecies before 1961 (see Art. 45.6.4 of the Code). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Valid as Gasterosteus crenobiontus Băcescu & Mayer 1956 but extinct -- (Kottelat 1997:165 [ref. 22952], Kottelat & Freyhof 2007:493 [ref. 29996]). Current status: Valid as Gasterosteus crenobiontus Băcescu & Mayer 1956. Gasterosteidae. Distribution: Freshwater springs, Lake Techirghiol, Romania [extinct]. Habitat: freshwater.

cuvieri, Gasterosteus Girard [C. F.] in Storer 1850:254, Pl. 7 (fig. 1) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Bras d'Or and Red Bay, Labrador, Canada. Non-types: MCZ 32812 (100), USNM 120288 [ex MCZ 32812] (6). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Smith 1986:277 [ref. 22186], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

dekayi, Gasterosteus Ayres [W. O.] 1855:48 [Proceedings of the California Academy of Sciences (Series 1) v. 1 (pt 1); ref. 159] Based on Gasterosteus biaculeatus of DeKay. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

dimidiatus, Gasterosteus Reinhardt [J. C. H.] 1837:114, 137, 193 [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 3691] Greenland. Syntypes: ZMUC. On pp. 32, 55 & 111 of separate. Reinhardt refers to Fauna Groenlandica Nr. 122 (Fabricius 1780:169-170 [ref. 17464]), which is Gasterosteus aculeatus in that work. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

elegans, Gasterosteus Blanchard [C. E.] 1866:234, Fig. 37 [Les poissons des eaux douces de la France; ref. 310] Between Cadillac and Langon, Gironde, southwestern France. Syntypes: MNHN 0000-6673 (2). Type catalog: Bertin & Estève 1950:38 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

gladiunculus, Gasterosteus Kendall [W. C.] 1896:623 [Proceedings of the United States National Museum v. 18 (no. 1089); ref. 11808] 4 miles southeast of Seguin Island, Maine, U.S.A. Holotype: USNM 47589. Paratypes: CAS-SU 4373 (3); USNM 47590-91 (1, 1), 125443-44 [ex USBF] (2, 1). Type catalog: Böhlke 1953:59 [ref. 12291]. •Synonym of Gasterosteus wheatlandi Putnam 1867 -- (Scott & Crossman 1998:671 [ref. 24570]). Current status: Synonym of Gasterosteus wheatlandi Putnam 1867. Gasterosteidae. Habitat: brackish, marine.

gymnurus, Gasterosteus Cuvier [G.] (ex Willughby) 1829:170 [Le Règne Animal (Edition 2) v. 2; ref. 995] England and northward; France. No types known. Based on Pisciculus aculeatus of Willughby 1686:341 [ref/ 30164]. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Valid as Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Kottelat & Freyhof 2007:494 [ref. 29996], Bianco & Delmastro 2011:6 [ref. 31648], Bianco 2013:9 [ref. 32897], Barbieri et al. 2015:96 [ref. 35029], Lorenzoni et al. 2019:252 [ref. 37108]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:972 [ref. 20662], Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2015:41 [ref. 35420], Denys et al. 2022:396 [ref. 40085], Çiçek et al. 2024:375 [ref. 40789]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish.

hologymnus, Gastrosteus Regan [C. T.] 1909:436 [Annals and Magazine of Natural History (Series 8) v. 4 (no. 23) (art. 50); ref. 14419] Rome, Italy. Syntypes: BMNH 1852.8.17.19-22 (4). Original genus should have been Gasterosteus. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:283 [ref. 7193], Lelek 1987:267 [ref. 22962], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

inopinatus, Gasterosteus Girard [C. F.] 1854:147 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] A fresh water lagoon about 1 mile back of Presidio, California, U.S.A. Syntypes: MCZ 6701 (1), MNHN 0000-0300 [ex USNM] (2). Type catalog: Bertin & Estève 1950:37 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

insculptus, Gasterosteus Richardson [J.] 1855:356, Pl. 25 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, western Greenland. Holotype (unique): whereabouts unknown. Not at BMNH. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

intermedius, Gasterosteus Girard [C. F.] 1856:135 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Cape Flattery, Washington, U.S.A. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

islandicus, Gasterosteus Sauvage [H.-E.] 1874:20, Pl. 1 (figs. 8-8c) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] Iceland. Holotype: MNHN 0000-5594 (1 of 3). Type catalog: Bertin & Estève 1950:38 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:283 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Valid as Gasterosteus islandicus Sauvage 1874 -- (Kottelat 1997:165 [ref. 22952], Kottelat & Freyhof 2007:496 [ref. 29996]). •Possible synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Denys et al. 2015:41 [ref. 35420]). Current status: Valid as Gasterosteus islandicus Sauvage 1874. Gasterosteidae. Distribution: Iceland. Habitat: freshwater.

japonicus, Gasterosteus williamsoni Franz [V.] 1910:19, Pl. 3 (fig. 10) [Abhandlungen der math.-phys. Klasse der K. Bayer Akademie der Wissenschaften v. 4 (Suppl.) (no. 1); ref. 1481] Misaki, Japan. Syntypes: ZSM [old collection] (2) destroyed in WWII. Type catalog: Neumann 2006:275 [ref. 28925]. •Permanently invalid, preoccupied by Gasterosteus japonicus Houttuyn 1782. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies microcephalus Girard 1854 -- (Okada 1961:612 [ref. 6363], Denys et al. 2022:398 [ref. 40085]). 1°Homonym. Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

johanseni, Gasterosteus bispinosus Cox [P.] 1923:147 [Canadian Field-Naturalist v. 37 (no. 8); ref. 15400] Surface in Westbay, Port au Port Bay, Newfoundland. Holotype (unique): whereabouts unknown. Gasterosteidae. Habitat: marine.

leiurus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:481, 487, Pl. 98 (fig. 4) [Histoire naturelle des poissons v. 4; ref. 998] France; Germany. Syntypes: MNHN 0000-7088 (3), 0000-7151 (7). •Synonym of Gasterosteus aculeatus Linnaeus 1758, but a valid subspecies -- (Kawanabe & Mizuno 1989:438 [ref. 25598], Nakabo 2002:513 [ref. 26001]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:972 [ref. 20662], Monod 1973:281 [ref. 7193], Reshetnikov et al. 1997:747 [ref. 24702] as a form, Reshetnikov & Savvaitove 1998:111 [ref. 23593], Scott & Crossman 1998:665 [ref. 24570], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

loricatus, Gasterosteus Reinhardt [J. C. H.] 1837:114, 193 [32, 111] [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 3691] Greenland. No types known. On pp. 32 & 111 of separate. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

messinicus, Gasterosteus aculeatus var. Stephanidis [A.] 1971:202, Fig. 10 [Biologia Gallo-Hellenica v. 3 (no. 2) (for 1970); ref. 7540] Not available, infrasubspecific; from Pamissos R. near Messini, Greece. •In the synonymy of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:398 [ref. 40085]). Infrasubspecific. Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

microcephalus, Gasterosteus Girard [C. F.] 1854:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] Tule Lake, San Joaquin Valley, California, U.S.A. Syntypes: ANSP 14275-87 (13), 14288-89 (2, missing); MNHN 0000-0302 [ex USNM] (1); UMMZ 146720 (4). Type catalog: Bertin & Estève 1950:36 [ref. 19576], Böhlke 1984:103 [ref. 13621]. •Valid as Gasterosteus microcephalus Girard 1854 -- (Herrera 1896:38 [ref. 39336], Okada 1961:612 [ref. 6363], Nakabo 2000:513 [ref. 25086], Mecklenburg et al. 2002:333 [ref. 25968]). •Synonym of Gaserosteus aculeatus Linnaeus 1758, but a valid subspecies microcephalus Girard 1854 -- (Kuroda 1953:277 [ref. 39545], Ida in Masuda et al. 1984:83 [ref. 6441], Swift et al. 1993:123 [ref. 22379], Castro-Aguirre et al. 1999:226 [ref. 24550], Ruiz-Campos et al. 2000:71 [ref. 24940], Moyle 2002:340 [ref. 26449], Miller 2006:320 [ref. 28615], Scharpf 2007:35 [ref. 30398]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:973 [ref. 20662], Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

nemausensis, Gasterosteus Crespon [J.] 1844:283 [Faune méridionale, ou description de tous les animaux vertébrés vivans et fosses v. 2; ref. 18056] Nîmes, France. No types known. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

neoboracensis, Gasterosteus DeKay [J. E.] 1842:66, Pl. 6 (fig. 17) [Zoology of New-York; ref. 1098] New York, U.S.A. No types known. Appeared first as name only in Dekay 1840:11 [ref. 17392]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Smith 1986:277 [ref. 22186], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

neustrianus, Gasterosteus Blanchard [C. E.] 1866:220, Fig. 28 [Les poissons des eaux douces de la France; ref. 310] Brooks in Harfleur and Gournay, France. Syntypes: MNHN 0000-7114 (1). Type catalog: Bertin & Estève 1950:37 [ref. 19576]. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

niger, Gasterosteus Cuvier [G.] 1829:170 [Le Règne Animal (Edition 2) v. 2; ref. 995] Terre Neuve, eastern Canada. Syntypes: MNHN 0000-3057 (2). Type catalog: Bertin & Estève 1950:36 [ref. 19576]. Available from footnote as, "G. niger ou biaculeatus, Mitchill. Trans. de New-Y., I, I, 10." Also appeared in Cuvier in Cuvier & Valenciennes 1829:503 [ref. 998]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

nipponicus, Gasterosteus Higuchi [M.], Sakai [H.] & Goto [A.] 2014:346, Figs. 1, 2a, 4 [Ichthyological Research v. 61 (no. 4); ref. 33316] Usujiri, Minami-kayabe, Hokkaido, Sea of Japan. Holotype: HUMZ 97486. Paratypes: Many at HUMZ, LBM, NSMT-P. Plus non-type specimens. •Valid as Gasterosteus nipponicus Higuchi, Sakai & Goto 2014 -- (Tohkairin et al. 2015:6 [ref. 35331], Dyldin & Orlov 2017:57 [ref. 35138], Dyldin et al. 2020:85, 110 [ref. 37569], Sonoyama et al. 2020:35 [ref. 37637], Dyldin et al. 2021:58 [ref. 38804], Dyldin & Orlov 2022:35 [ref. 38972]). Current status: Valid as Gasterosteus nipponicus Higuchi, Sakai & Goto 2014. Gasterosteidae. Distribution: Northwestern Pacific: Sea of Japan, Sea of Okhotsk, south to Korea and Sakhalin Island (Russia). Habitat: freshwater, brackish, marine.

noveboracensis, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:502, Pl. 98 (fig. 3) [Histoire naturelle des poissons v. 4; ref. 998] New York, U.S.A. Holotype (unique): MNHN 0000-7074. Type catalog: Bertin & Estève 1950:35 [ref. 19576]. Appeared first as name only in Cuvier 1829:170 [ref. 995]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

obolarius, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:500 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

plebeius, Gasterosteus Girard [C. F.] 1854:147 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] Salt marshes about Presidio, California, U.S.A. Syntypes and/or Girard specimens: FMNH 6038 (4) Petaluma, MCZ 6706 (6) Petaluma, California, MNHN 0298 [ex USNM] (8) Petaluma; ?UMMZ 146720 or 146721 (1). Type catalog: Bertin & Estève 1950:36-37 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

ponticus, Gasterosteus Nordmann [A. von] 1840:380, 613, Pl. 3 (fig. 11) [Voyage dans la Russie méridionale et la Crimée; ref. 3209] Tauria, Black Sea. Syntypes: MNHN 0000-7098 (3), NMW 77244 (4). Type catalog: Bertin & Estève 1950:36 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:972 [ref. 20662], Monod 1973:281 [ref. 7193], Kottelat 1997:164 [ref. 22952], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Parin et al. 2014:203 [ref. 33547], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

pugetti, Gasterosteus Girard [C. F.] 1856:135 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Fort Steilacoom, Puget Sound, Washington, U.S.A. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

quadrispinosa, Gasterosteus Crespon [J.] 1844:283 [Faune méridionale, ou description de tous les animaux vertébrés vivans et fosses v. 2; ref. 18056] Vistre River and ditches around Nîmes, France. Syntypes: whereabouts unknown. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

santaeannae, Gasterosteus Regan [C. T.] 1909:437 [Annals and Magazine of Natural History (Series 8) v. 4 (no. 23) (art. 50); ref. 14419] Colton, Santa Ana River, California, U.S.A. Syntypes: BMNH 1894.2.2.15-17 (3). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Denys et al. 2022:398 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

semiarmatus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:493 [Histoire naturelle des poissons v. 4; ref. 998] Le Havre and Braie River near Abbeville, France. Syntypes: MNHN 0000-2347 (1), 0000-2348 (1), 0000-7078 (3). Type catalog: Bertin & Estève 1950:34-35 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Lelek 1987:267 [ref. 22962], Reshetnikov et al. 1997:747 [ref. 24702] as a form, Reshetnikov & Savvaitove 1998:111 [ref. 23593], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

semiloricatus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:494 [Histoire naturelle des poissons v. 4; ref. 998] Somme River, environs de Paris, Oise, Rochelle, Normandie, Caen, Hable d'Ault, Treport, Berlin. Syntypes: MNHN 0000-2347 (1). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

serratus, Gasterosteus Ayres [W. O.] 1855:[2] col. 6 [The Pacific [newspaper] v. 4 (nos 7-22); ref. 13428] San Francisco, California, U.S.A. Holotype (unique): apparently lost. Non-types: MNHN 0299 [ex USNM] (1). Type catalog: Bertin & Estève 1950:37 [ref. 19576]. Appeared first as above on 4 May; then in Ayres 1855:47 [ref. 159]. Monod 1973:282 [ref. 7193] lists a syntype at MNHN. •Apparently preoccupied by Gasterosteus serratus Forster 1794. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). 1°Homonym. Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

spinulosus, Gasterosteus Yarrell [W.] (ex Jenyns) 1835:350 [A manual of British vertebrate animals; ref. 18335] Edinburgh, Scotland. No types known. Also appeared in Yarrell 1836:83, fig. [ref. 4812], earliest not researched. •Tentatively a synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Kottelat 1997:164 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856] as spinosus, Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

suppositus, Gasterosteus Sauvage [H.-E.] 1874:11 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] New York, U.S.A. Based on Gasterosteus noveboracensis of Dekay 1842:66 [ref. 1098]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:283 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

teraculeatus, Gasterosteus Lacepède [B. G. E.] 1801:295, 296 [Histoire naturelle des poissons (Lacepède) v. 3; ref. 2710] Unneeded new name for Gasterosteus aculeatus Linnaeus 1758. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Kottelat 1997:163 [ref. 22952], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

tetracanthus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:499 [Histoire naturelle des poissons v. 4; ref. 998] Florence, Italy. Holotype (unique): MNHN 0000-1821. Type catalog: Bertin & Estève 1950:35 [ref. 19576]. Appeared first as name only in Cuvier 1829:170 [ref. 995]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856]). •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

texanus, Gasterosteus Sauvage [H.-E.] 1874:15, Pl. 1 (fig. 5-5a) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] Texas, U.S.A. Holotype (unique): MNHN 0000-7076. Type catalog: Bertin & Estève 1950:38 [ref. 19576]. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

trachurus, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:481, 487, Pl. 98 (fig. 1) [Histoire naturelle des poissons v. 4; ref. 998] France; Germany. Syntypes: ?NMW 6918 (1). •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Berg 1949:972 [ref. 20662], Monod 1973:281 [ref. 7193], Lelek 1987:267 [ref. 22962], Kottelat 1997:164 [ref. 22952], Reshetnikov et al. 1997:747 [ref. 24702] as a form, Reshetnikov & Savvaitove 1998:111 [ref. 23593], Scott & Crossman 1998:665 [ref. 24570], Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:396 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.

wheatlandi, Gasterosteus Putnam [F. W.] 1867:4 [Proceedings of the Essex Institute v. 5 (1866-67); ref. 18577] Nahant, Massachusetts, U.S.A. Syntypes: whereabouts unknown. •Valid as Gasterosteus wheatlandi Putnam 1867 -- (Lee et al. 1980:565 [ref. 22416], Robins & Ray 1986:122 [ref. 23100], Scott & Scott 1988:343 [ref. 25518], McAllister 1990:217 [ref. 14674], Scott & Crossman 1998:669 [ref. 24570], Krueger in Collette & Klein-MacPhee 2002:316 [ref. 26158], Nelson et al. 2004:111 [ref. 27807], Page et al. 2013:113 [ref. 32708]). Current status: Valid as Gasterosteus wheatlandi Putnam 1867. Gasterosteidae. Distribution: Eastern North America. Habitat: brackish, marine.

williamsoni, Gasterosteus Girard [C. F.] 1854:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] Upper part of Santa Clara River, Soledad Canyon, Los Angeles County, California, U.S.A. Syntypes: MNHN 0000-0303 [ex USNM] (1), USNM 340 (now 1). Type catalog: Bertin & Estève 1950:36 [ref. 19576]. For information on type locality see Miller 1960 [ref. 21987]. •Synonym of Gasterosteus aculeatus Linnaeus 1758, subspecies aculeatus -- (Okada 1961:605 [ref. 6363]). •Synonym of Gaserosteus aculeatus Linnaeus 1758, but a valid subspecies -- (Swift et al. 1993:125 [ref. 22379], Moyle 2002:340 [ref. 26449], Scharpf 2007:35 [ref. 30398]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:282 [ref. 7193], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.