An electronic journal published by the California Academy of Sciences
ISSN 1545-150X [CD-ROM]
View Annotated Checklists arranged
Alphabetical List of Fish Families and Subfamilies
Click on a highlighted family name to view or download the checklist in postscript document format [PDF]
Introduction to the series Annotated Checklists of Fishes
A
Abyssocottidae
Acanthoclininae (spiny basslets)
Acanthuridae (surgeonfishes and unicornfishes)
Acestrorhynchidae
Achiridae
Achiropsettidae (southern or armless flounders)
Acipenseridae (sturgeons)
Acropomatidae (lanternbellies)
Adrianichthyidae
Adrianichthyinae (adrianichthyids)
Ageneiosidae (bottlenose or barbelless catfishes)
Agonidae (poachers, starsnouts, rockheads and alligator fishes)
Agoninae
Akysidae (stream catfishes)
Albulidae (bonefishes)
Albulinae
Alepisauridae (lancetfishes)
Alepocephalidae (slickheads)
Alestiidae (African tetras)
Alopiidae (thresher sharks)
Amarsipidae (amarsipids)
Ambassidae
Amblycipitidae (torrent catfishes)
Amblyopinae
Amblyopsidae (cavefishes)
Amiidae (bowfins)
Ammodytidae (sand lances)
Amphiliidae
Amphiliidae (loach catfishes)
Amphiliinae
Anabantidae (climbing gouramies)
Anablepidae
Anablepinae (four-eyed fishes)
Anacanthobatidae (smooth skates)
Anarhichadidae (wolffishes)
Anguillidae (freshwater eels)
Anisochrominae
Anomalopidae (lanterneye fishes)
Anoplagoninae
Anoplogastridae (fangtooths)
Anoplopomatidae (sablefishes)
Anostomidae
Anostominae
Anotopteridae (daggertooths)
Antennariidae (frogfishes)
Anthiinae
Aphanopodinae
Aphredoderidae (pirate perches)
Aphyonidae (aphyonids)
Apistidae
Aploactinidae (velvetfishes)
Aplocheilichthyinae
Aplocheilidae (Old World rivulines)
Aplochitoninae
Aplodactylidae
Apogonidae (cardinalfishes)
Apogoninae
Apteronotidae (ghost knifefishes)
Aracaninae
Arapaimidae (arapaimas)
Argentinidae (argentines)
Ariidae (sea catfishes)
Ariommatidae (ariommatids)
Arripidae (Australian salmons)
Artedidraconidae
Aspredinidae (banjo catfishes)
Aspredininae
Astroblepidae (climbing catfishes)
Astronesthinae (snaggletooths)
Ateleopodidae (jellynose fishes)
Atherinidae (silversides)
Atherininae
Atherinopsinae
Atherioninae
Auchenipteridae (auchenipterid catfishes)
Aulopidae (aulopus)
Aulorhynchidae (tubesnouts)
Aulostomidae (trumpetfishes)
B
Badinae
Bagridae (bagrid catfishes)
Balistidae (triggerfishes)
Balitoridae (hillstream and river loaches)
Balitorinae (flat loaches)
Banjosidae
Barbourisiidae (velvet whalefishes)
Bathyagoninae
Bathyclupeidae (bathyclupeids)
Bathydraconidae (Antarctic dragonfishes)
Bathygadinae
Bathylagidae (deepsea smelts)
Bathylutichthyidae
Bathymasteridae (ronquils)
Bathymyrinae
Bathysaurinae
Bathysauroididae
Bathysauropsidae
Batrachoididae (toadfishes)
Bedotiidae
Belonidae (needlefishes)
Belontiinae
Bembridae (deepwater flatheads)
Bembropinae
Berycidae (berycids or alfonsinos)
Blenniidae (blennies)
Bothidae (lefteye flounders)
Bothragoninae (rockheads)
Botiinae
Bovichtidae (bovichtids)
Brachaeluridae (blind sharks)
Brachionichthyidae (warty anglerfishes)
Brachyopsinae
Bramidae (pomfrets)
Bregmacerotidae (codlets)
Brosmophycinae
Brotulinae
Brotulotaeniinae
Bunocephalinae
Butinae
Bythitidae (livebearing brotulas)
Bythitinae
C
Caesionidae (fusiliers)
Callanthiidae
Callichthyidae (callichthyid armored catfishes)
Callionymidae (dragonets)
Callorhinchidae (plownose chimaeras)
Caproidae
Caracanthidae (coral crouchers)
Carangidae (jacks, amberjacks, pompanos)
Carapidae
Carapinae (pearlfishes)
Carcharhinidae (requiem sharks)
Caristiidae (veilfins or manefishes)
Catostomidae (suckers)
Caulophrynidae
Centracanthidae
Centrarchidae (sunfishes and freshwater basses)
Centriscidae
Centriscinae (shrimpfishes)
Centrogeniidae
Centrolophidae (medusafishes)
Centrophoridae
Centrophrynidae
Centropomidae
Centropominae (snooks)
Cepolidae (bandfishes)
Ceratiidae (seadevils)
Ceratodontidae (Australian lungfishes)
Cetomimidae (whalefishes)
Cetopsidae (whalelike catfishes)
Cetorhinidae (basking sharks)
Chacidae (squarehead or angler catfishes)
Chaenopsidae (pikeblennies)
Chaetodontidae (butterflyfishes)
Champsodontidae (gapers)
Chanidae (milkfishes)
Channichthyidae (crocodile icefishes)
Channidae (snakeheads)
Characidae (characins)
Characidiinae
Chaudhuriidae
Chauliodontinae (viperfishes)
Chaunacidae (sea toads)
Cheilobranchinae
Cheilodactylidae (fingerfins or morwongs)
Chiasmodontidae (swallowers)
Chilodontinae
Chimaeridae (shortnose chimaeras or ratfishes)
Chirocentridae (wolf herrings)
Chirolophinae
Chironemidae (kelpfishes)
Chlamydoselachidae (frill sharks)
Chlopsidae (false morays)
Chlorophthalmidae (greeneyes)
Choridactylinae
Cichlidae (cichlids)
Cirrhitidae (hawkfishes)
Citharidae
Citharinidae
Citharininae
Clariidae (airbreathing or labyrinth catfishes)
Clinidae (clinids and klipfishes)
Clupeidae (herrings, shads, sardines and allies)
Cobitidae (loaches)
Cobitinae
Coiidae (tigerperches)
Colocongridae
Comephoridae (Baikal oilfishes)
Congiopodidae (horsefishes)
Congridae (conger eels)
Congrinae
Congrogadinae (eelblennies)
Coregoninae (whitefishes)
Coryphaenidae (dolphins or dolphinfishes)
Cottidae (sculpins)
Cottocomephoridae
Cranoglanididae (armorhead catfishes)
Creediidae (sand burrowers)
Cryptacanthodidae (wrymouths)
Ctenoluciidae (pike-characids)
Cubanichthyinae
Curimatidae
Curimatinae (curimatas)
Cyclopteridae (lumpsuckers)
Cyematidae (bobtail eels)
Cynodontidae
Cynodontinae
Cynoglossidae (tonguefishes)
Cynoglossinae
Cyprinidae (minnows and carps)
Cyprinodontidae (killifishes)
Cyprinodontinae
Cyttidae
Cyttopsinae
D
Dactylopteridae (helmet gurnards)
Dactyloscopidae (sand stargazers)
Dalatiidae
Dalatiinae
Dasyatidae (stingrays)
Dentatherinidae
Denticipitidae (denticle herrings)
Derichthyidae (longneck eels)
Diceratiidae
Dichistiidae (galjoens)
Dinolestidae
Dinopercidae (cavebasses)
Diodontidae (porcupinefishes and burrfishes)
Diplomystidae (diplomystid catfishes)
Diretmidae (spinyfins)
Distichodontinae
Doradidae (thorny catfishes)
Doumeinae
Draconettidae (draconettids)
Drepaneidae (sicklefishes)
E
Echeneidae (remoras and sharksuckers)
Echinorhinidae (bramble sharks)
Elassomatidae (pygmy sunfishes)
Electrophoridae (electric knifefishes)
Eleginopidae
Eleotridae (sleepers)
Eleotrinae
Elopidae (tenpounders and ladyfishes)
Embiotocidae (surfperches)
Emmelichthyidae (rovers)
Empetrichthyinae
Engraulidae (anchovies)
Enoplosidae (oldwives)
Ephippidae (spadefishes, batfishes and scats)
Epigonidae
Epinephelinae (groupers)
Eptatretinae
Erethistidae
Ereuniidae
Erythrinidae (trahiras)
Esocidae (pikes)
Etmopterinae
Euclichthyidae (eucla cods)
Eurypharyngidae (umbrellamouth gulpers)
Evermannellidae (sabertooth fishes)
Exocoetidae (flyingfishes)
F
Fistulariidae (cornetfishes)
Fluviphylacinae
Fundulidae
G
Gadidae (cods and haddocks)
Galaxiidae (galaxiids)
Galaxiinae
Galeorhininae
Gasteropelecidae (freshwater hatchetfishes)
Gasterosteidae (sticklebacks)
Gempylidae (snake mackerels)
Geotriinae
Gerreidae (mojarras)
Gibberichthyidae (gibberfishes)
Gigantactinidae (whipnoses)
Giganturidae (telescopefishes)
Ginglymostomatidae (nurse sharks)
Girellinae (nibblers)
Glandulocaudinae
Glaucosomatidae
Gnathanacanthidae (red velvetfishes)
Gobiesocidae (clingfishes)
Gobiesocinae
Gobiidae (gobies)
Gobiinae (gobies)
Gobionellinae
Gonorynchidae
Gonostomatidae (bristlemouths)
Goodeidae (goodeids)
Goodeinae
Grammatidae (basslets)
Grammicolepididae
Grammicolepidinae
Grammistinae (soapfishes)
Gymnarchidae
Gymnelinae
Gymnotidae (nakedback knifefishes)
Gymnuridae (butterfly rays)
Gyrinocheilidae (algae eaters)
H
Haemulidae (grunts)
Halosauridae (halosaurs)
Harpadontinae (Bombay ducks)
Harpagiferidae (plunderfishes)
Helogeneidae (helogeneid catfishes)
Helostomatidae (kissing gouramies)
Hemerocoetinae
Hemigaleidae (weasel sharks)
Hemiodontidae
Hemiodontinae
Hemiramphidae (halfbeaks)
Hemiramphinae
Hemiscylliidae (bamboo sharks)
Hemitripteridae (sailfin sculpins or sea ravens)
Hepsetidae
Heterenchelyidae
Heterocongrinae (garden eels)
Heterodontidae (bullhead and horn sharks)
Heteropneustidae (airsac catfishes)
Hexagrammidae (greenlings)
Hexagramminae (greenlings)
Hexanchidae (cow sharks)
Hexatrygonidae (sixgill stingrays)
Himantolophidae (footballfishes)
Hiodontidae (mooneyes)
Hippocampinae (seahorses)
Hispidoberycidae (hispidoberycids)
Holocentridae (squirrelfishes and soldierfishes)
Holocentrinae (squirrelfishes)
Hoplichthyidae (spiny flatheads)
Hoplomyzontinae
Horaichthyinae
Hypninae
Hypophthalmidae (loweye catfishes)
Hypopomidae
Hypoptychidae (sand-eels)
Hypsagoninae
I
Icosteidae (ragfishes)
Ictaluridae (North American freshwater catfishes)
Idiacanthinae (black dragonfishes)
Ilyophinae (arrowtooth eels)
Indostomidae
Inermiidae (bonnetmouths)
Ipnopidae
Istiophoridae (billfishes and marlins)
K
Kneriidae
Kraemeriidae (sand darts)
Kuhliidae (flagtails or aholeholes)
Kurtidae (nurseryfishes)
Kyphosidae (sea chubs)
Kyphosinae (rudderfishes)
L
Labridae (wrasses)
Labrisomidae (labrisomids)
Lactariidae (false trevallies)
Lamnidae (mackerel sharks)
Lampridae (opahs)
Lateolabracidae (Asian seaperches)
Latilinae (tilefishes)
Latimeriidae (coelacanths)
Latinae
Latridae (trumpeters)
Lebiasinidae
Lebiasininae
Leiognathidae (ponyfishes or slipmouths)
Lepidogalaxiidae (salamanderfishes)
Lepidopinae
Lepidosirenidae (South American lungfishes)
Lepisosteidae (gars)
Leptobramidae (beachsalmon)
Leptochariidae (barbeled houndsharks)
Leptochilichthyidae (leptochilichthyids)
Leptoglanidinae
Leptoscopidae
Lethrinidae (emperors)
Linophrynidae
Liopropomatinae
Liparidae (snailfishes)
Lobotidae (tripletails)
Lophichthyinae
Lophiidae (goosefishes or monkfishes)
Lophotidae (crestfishes)
Loricariidae (suckermouth armored catfishes)
Lotidae (hakes and burbots)
Luciocephalinae
Lumpeninae
Lutjanidae (snappers)
Luvaridae (luvars)
Lycodinae
Lycozoarcinae
M
Macropodinae
Macroramphosinae (snipefishes)
Macrouridae (grenadiers or rattails)
Macrourinae
Macrouroidinae (southern hakes)
Macrurocyttinae
Malacanthidae (tilefishes)
Malacanthinae (sand tilefishes)
Malacosteinae (loosejaws)
Malapteruridae (electric catfishes)
Mastacembelidae (spiny-eels)
Maurolicinae
Megachasmidae (megamouth sharks)
Megalomycteridae (largenose fishes)
Megalopidae (tarpons)
Melamphaidae (bigscales)
Melanocetidae
Melanonidae (pelagic cods)
Melanostomiinae (scaleless black dragonfishes)
Melanotaeniidae (rainbowfishes)
Menidae (moonfishes)
Menidiinae
Merlucciidae (merluccid hakes)
Merlucciinae
Microcanthinae
Microdesmidae
Microdesminae (wormfishes)
Microstomatidae
Minoinae
Mirapinnidae (hairyfishes and ribbontails)
Mitsukurinidae (goblin sharks)
Mobulinae (manta and devil rays)
Mochokidae (squeakers and unside-down catfishes)
Molidae (molas or ocean sunfishes)
Monacanthidae (filefishes)
Monocentridae (pinecone fishes)
Monodactylidae (moonies)
Monognathidae (monognathids)
Mordaciinae
Moridae (deepsea cods)
Moringuidae (spaghetti eels)
Mormyridae (elephantfishes)
Moronidae
Mugilidae (mullets)
Mullidae (goatfishes)
Muraenesocidae (pike conger eels)
Muraenidae (morays)
Muraeninae
Muraenolepididae (eel cods)
Myctophidae (lanternfishes)
Myliobatidae
Myliobatinae (eagle rays)
Myripristinae (soldierfishes)
Myrocongridae
Myrophinae
Myxinidae (Hagfishes)
Myxininae
N
Nandidae (leaffishes)
Nandinae
Narcinidae
Narcininae
Narkinae
Nemacheilinae
Nematistiidae (roosterfishes)
Nemichthyidae (snipe eels)
Nemipteridae (threadfin breams and spinycheeks)
Neobythitinae
Neoceratiidae
Neoscopelidae (blackchins)
Neosebastidae
Neozoarcinae
Nettastomatidae (duckbill eels)
Nomeidae (flotsamfishes or driftfishes)
Normanichthyidae
Notacanthidae (spiny eels)
Notocheiridae
Notograptidae
Notopteridae (knifefishes or featherbacks)
Notosudidae (paperbones or waryfishes)
Nototheniidae (notothens)
O
Odacidae
Odontaspididae (sand tiger sharks)
Odontobutidae
Ogcocephalidae (batfishes)
Olyridae (olyrid catfishes)
Omosudidae (omosudids)
Oneirodidae
Ophichthidae (snake eels and worm eels)
Ophichthinae
Ophidiidae (cuskeels)
Ophidiinae
Ophiodontinae (lingcods)
Opisthocentrinae
Opisthoproctidae (spookfishes or barreleyes)
Opistognathidae (jawfishes)
Oplegnathidae (knifejaws)
Orectolobidae (wobbegons)
Oreosomatidae
Oreosomatinae
Oryziinae (medakas or ricefishes)
Osmeridae (smelts)
Osphronemidae (giant gouramies)
Osphroneminae
Osteoglossidae (bonytongues)
Ostraciidae (boxfishes)
Ostraciinae
Ostracoberycidae
Oxudercinae (mudskippers and allies)
Oxylebiinae (painted greenlings)
Oxynotinae
Oxyzygonectinae
P
Pangasiidae (pangasid catfishes)
Pantodontidae
Parabembridae
Parabrotulidae (false brotulas)
Parakysidae (parakysid catfishes)
Paralepididae (barracudinas)
Paralichthodinae
Paralichthyidae
Parascorpididae (jutjaws)
Parascylliidae (collared carpet sharks)
Paraulopidae
Parazenidae
Parazeninae
Parodontinae
Pataecidae (prowfishes)
Pegasidae (seamoths)
Pempheridae (sweepers)
Pentacerotidae (armorheads)
Percichthyidae (temperate basses)
Percidae (perches and darters)
Percophidae (duckbills)
Percophinae
Percopsidae (trout-perches)
Peristediidae (armored gurnards)
Petromyzontidae (lampreys)
Petromyzontinae
Phallostethidae
Pholidae (gunnels)
Pholidichthyidae (convict-blennies)
Phosichthyidae (lightfishes)
Phractolaemidae
Phycidae (phycid hakes)
Pimelodidae (long-whiskered catfishes)
Pinguipedidae (sandperches)
Platycephalidae (flatheads)
Platyrhininae (thornbacks)
Platytroctidae (tubeshoulders)
Plecoglossidae (ayu fishes)
Plectrogenidae
Plesiobatidae (deepwater stingrays)
Plesiopidae (roundheads)
Plesiopinae
Pleurogramminae (Atka mackerels)
Pleuronectidae (righteye flounders)
Pleuronectinae
Plotosidae (eeltail catfishes)
Poeciliidae (poeciliids)
Poeciliinae (livebearers)
Poecilopsettinae
Polymixiidae (beardfishes)
Polynemidae (threadfins)
Polyodontidae (paddlefishes)
Polyprionidae (wreckfishes)
Polypteridae (bichirs)
Pomacanthidae (angelfishes)
Pomacentridae (damselfishes and anemonefishes)
Pomatomidae (bluefishes)
Potamotrygonidae (river stingrays)
Priacanthidae (bigeyes)
Pristidae (sawfishes)
Pristiophoridae (saw sharks)
Pristolepidinae
Prochilodontinae
Profundulidae
Proscylliidae (finback catsharks)
Protopteridae (African lungfishes)
Prototroctinae (southern graylings)
Psettodidae (spiny turbots)
Pseudaminae
Pseudaphritidae
Pseudocarchariidae (crocodile sharks)
Pseudochromidae (dottybacks)
Pseudochrominae
Pseudocyttinae
Pseudomugilidae (blue eyes)
Pseudoplesiopinae
Pseudotriakidae (false catsharks)
Pseudotrichonotidae
Psilorhynchidae
Psychrolutidae (fatheads)
Ptereleotrinae
Pteroinae (turkeyfishes and lionfishes)
Pterothrissinae
Ptilichthyidae (quillfishes)
Pyramodontinae (pyramodontines)
Pyrrhulininae
R
Rachycentridae (cobias)
Radiicephalidae (tapertails)
Rajidae (skates)
Regalecidae (oarfishes)
Retropinnidae (New Zealand smelts)
Retropinninae (southern smelts)
Rhamphichthyidae (sand knifefishes)
Rhamphocottidae (horsehead sculpins or grunt sculpins)
Rhincodontidae (whale sharks)
Rhininae
Rhinobatidae (guitarfishes)
Rhinobatinae
Rhinochimaeridae (longnose chimaeras)
Rhinopterinae (cownose rays)
Rhombosoleinae
Rhyacichthyidae (loach gobies)
Rhynchobatinae
Rivulidae (New World rivulines)
Roestinae
Rondeletiidae (redmouth whalefishes)
S
Saccopharyngidae (swallowers or whiptail gulpers)
Salangidae (icefishes or noodlefishes)
Salmonidae (salmonids)
Salmoninae (salmons, trouts, chars and allies)
Samaridae
Scaridae (parrotfishes)
Scarinae
Scatophagidae (scats)
Schilbeidae (schilbeid catfishes)
Schindleriidae
Sciaenidae (croakers and drums)
Scoloplacidae (spiny dwarf catfishes)
Scomberesocidae (sauries)
Scombridae (mackerels, tunas and bonitos)
Scombrolabracidae (black mackerels)
Scombropidae (gnomefishes)
Scopelarchidae (pearleyes)
Scophthalmidae (scophthalmids)
Scorpaenidae (scorpionfishes and allies)
Scorpaeninae (scorpionfishes)
Scorpidinae (halfmoons)
Scyliorhinidae (cat sharks)
Scytalinidae (graveldivers)
Sebastidae
Sebastinae (rockfishes and rockcods)
Sebastolobinae (thornyheads)
Serranidae (sea basses and groupers)
Serraninae
Serrasalminae (piranhas and allies)
Serrivomeridae (sawtooth eels)
Setarchidae
Sicydiinae
Siganidae (rabbitfishes)
Sillaginidae (sillagos)
Siluridae (sheatfishes)
Simenchelyinae (pugnose eels)
Sisoridae (sisorid catfishes)
Soleidae (soles)
Solenostomidae (ghost pipefishes)
Somniosinae (sleeper sharks)
Sparidae (porgys and seabreams)
Sparisomatinae
Sphyraenidae (barracudas)
Sphyrnidae (hammerhead sharks)
Squalidae (dogfish sharks)
Squatinidae (angel sharks)
Stegostomatidae (zebra sharks)
Steindachneriinae
Stephanoberycidae (pricklefishes)
Sternoptychidae
Sternoptychinae (marine hatchetfishes)
Sternopygidae (glass knifefishes)
Stethaprioninae
Stichaeidae (pricklebacks)
Stichaeinae
Stomiidae
Stomiinae (saly dragonfishes)
Stromateidae (butterfishes)
Stylephoridae (tube-eyes or thread-tails)
Sundasalangidae (Sundaland noodlefishes)
Symphurinae
Symphysanodontidae
Synanceiidae
Synanceiinae (stonefishes and allies)
Synaphobranchidae
Synaphobranchinae (cutthroat eels)
Synbranchidae (swamp-eels)
Syngnathidae
Syngnathinae (pipefishes)
Synodontidae
Synodontinae (lizardfishes)
T
Telmatherinidae
Terapontidae (grunters or tigerfishes)
Tetrabrachiidae
Tetragonuridae (squaretails)
Tetraodontidae (puffers)
Tetrarogidae (waspfishes)
Thaumatichthyidae
Thymallinae (graylings)
Torpedinidae (electric rays)
Torpedininae
Toxotidae (archerfishes)
Trachichthyidae (slimeheads)
Trachinidae (weeverfishes)
Trachipteridae (ribbonfishes)
Trachyrincinae
Triacanthidae (triplespines)
Triacanthodidae (spikefishes)
Triakidae (houndsharks)
Triakinae
Trichiuridae (cutlassfishes and scabbardfishes)
Trichiurinae
Trichodontidae (sandfishes)
Trichomycteridae (parasitic catfishes and candirus)
Trichonotidae (sand divers)
Triglidae (searobins)
Triodontidae (threetooth puffers)
Tripterygiidae (threefin blennies)
U
Umbridae (mudminnows)
Uranoscopidae (stargazers)
Urolophidae (round stingrays)
Uropterygiinae
V
Valenciidae
Veliferidae (velifers)
X
Xenisthmidae
Xiphiidae (swordfishes)
Xiphisterinae
Z
Zanclidae (moorish idols)
Zaniolepidinae (combfishes)
Zaproridae (prowfishes)
Zeidae
Zenarchopterinae
Zenionidae
Zoarcidae (eelpouts)
Zoarcinae
This material is based upon work supported by the National Science Foundation under Grant No. 0208690.
Comments and questions related to this information:
Comments and questions related to web server:
Page created by
URL: http://www.calacademy.org/research/ichthyology/annotated/index.html
Last updated: 3-September-2003 / WJP
About the Department |
General information and a brief history of the Department of Ichthyology.
Ichthyology Staff |
Ichthyology curators, collection managers, curatorial assistants, and research associates.
The Catalog of Fishes |
The authoritative reference for taxonomic fish names, featuring a searchable on-line database.
Collection Database |
Search our on-line database or find out more about the collection.
Primary Types Imagebase |
Search for photographs, radiographs, and illustrations of our primary types.
Annotated Checklists |
This series serves to provide the most up-to-date taxonomic treatment of the families of recent fishes.
Funding Opportunities |
Financial aid for research in the Department of Ichthyology.
Newsletter of Systematic |
Browse past issues of the newsletter.
Contact Info |
For general inquiries about the Department of Ichthyology please contact:
Ichthyology Administrator
415-379-5277